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A new name for the oviraptorid dinosaur “Ingeniayanshini
(Barsbold, 1981; preoccupied by Gerlach, 1957)



The genus name of the small oviraptorid dinosaur Ingenia yanshini is preoccupied by the tripyloidid nematode Ingenia mirabilis, thus making the former a junior homonym of the latter.  Although “Ingeniayanshini is sympatric with Conchoraptor gracilis, it is distinguished from Conchoraptor by proportions of the manus.  It also differs appreciably from the ingeniines Heyuannia huangi and Nemegtomaia barsboldi in manual and pelvic proportions.  “Ingeniayanshini is not referable to any other taxon, and is renamed Ajancingenia yanshini gen. nov.  Several specimens originally referred to this species have subsequently been transferred to new genera, and the taxonomic assignment of material referred to Ajancingenia yanshini gen. nov. is reassessed.


Theropoda, Oviraptorosauria, Ingenia, Ajancingenia, Barun Goyot Formation

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Azuma, Y. (2005) The Flying Dinosaurs. Fukui Prefectural Dinosaur Museum, Katsuyama, 118 pp.

Barsbold, R. (1977) [On the evolution of carnivorous dinosaurs]. Joint Soviet-Mongolian Paleontological Expedition Transactions, 4, 48–56. [in Russian with English summary]

Barsbold, R. (1981) [Toothless carnivorous dinosaurs of Mongolia]. Joint Soviet-Mongolian Paleontological Expedition Transactions, 15, 28–39. [in Russian]

Barsbold, R. (1986) Raubdinosaurier Oviraptoren. In: Vorobyeva, E.I. (Ed.), Herpetological Studies in the Mongolian People’s Republic. Akademia Nauk SSSR Institut Evolyucionnoy Morfologii i Ekologii Zhivotnikhim. Moskva: A.M. Severtsova, pp. 210–223. [in Russian, with German summary]

Barsbold, R., Maryańska, T. & Osmólska, H. (1990) Oviraptorosauria. In: Weishampel, D.B., Dodson, P. & Osmólska, H. (Eds.), The Dinosauria 1st ed. University of California Press, Berkeley, pp. 249–258.

Clark, J.M., Norell, M.A. & Barsbold, R. (2001) Two new oviraptorids (Theropoda: Oviraptorosauria), Upper Cretaceous Djadokhta Formation, Ukhaa Tolgod, Mongolia. Journal of Vertebrate Paleontology, 21, 209–213.[0209:tnotou];2

Choiniere, J.N., Clark, J.M., Forster, C.A., Norell, M.A., Eberth, D.A., Erickson, G.M., Chu, H. & Xu, X. (2013) A juvenile specimen of a new coelurosaur (Dinosauria: Theropoda) from the Middle-Late Jurassic Shishugou Formation of Xinjiang, People’s Republic of China. Journal of Systematic Paleontology, 2013, 1–39.

Currie, P.J. (2002) Report on fieldwork in Mongolia, September 2001. In: Alberta Paleontological Society, sixth annual symposium, “Fossils 2002”. Presented by Alberta Paleontological Society, in conjunction with Canadian Society of Petroleum Geologists, Paleontological Division and Department of Earth Sciences, Mount Royal College, pp. 8–12.

Dashzeveg, D., Novacek, M.J., Norell, M.A., Clark, J.M., Chiappe, L.M., Davidson, M.C., McKenna, L., Dingus, C., Swisher, C. & Perle, A. (1995) Unusual preservation in a new vertebrate assemblage from the Late Cretaceous of Mongolia. Nature, 374, 446–449.

Dingus, L., Gaffney, E.S., Norell, M.A. & Sampson, S. (1995) The Halls of Dinosaurs—A Guide to Saurischians and Ornithischians. American Museum of Natural History, New York, 100 pp.

Fanti, F., Currie, P.J. & Badamgarav, D. (2012) New Specimens of Nemegtomaia from the Baruungoyot and Nemegt Formations (Late Cretaceous) of Mongolia. PLoS ONE, 7(2), e31330.

Fara, E. (2001) Khermeen-Tsav locality (SMPE) (Cretaceous of Mongolia). Available from: (Accessed 5 August 2013)

Gerlach, S.A. (1957) Die Nematodenfauna des Sandstrandes an der Küste von Mittelbrasilien (Brasilianische Meeres-Nematoden IV). Mitteilungen aus dem Museum für Naturkunde in Berlin. Zoologisches Museum und Institut für Spezielle Zoologie (Berlin), 33, 411–459.

Gubin, Y.M. (1999) [Gobiatids (Anura) from the Upper Cretaceous locality Khermeen-Tsav (Gobi Desert, Mongolia)]. Paleontologicheskii Zhurnal, 1, 76–87. [in Russian]

Kundrát, M. (2007) Avian-like attributes of a virtual brain model of the oviraptorid theropod Conchoraptor gracilis. Naturwissenschaften, 94, 499–504.

Longrich, N.R., Currie, P.J. & Dong, Z.-M. (2010) A new oviraptorid (Dinosauria: Theropoda) from the Upper Cretaceous of Bayan Mandahu, Inner Mongolia. Paleontology, 53, 945–960.

Longrich, N.R., Barnes, K., Clark, S. & Millar, L. (2013) Caenagnathidae from the Upper Campanian Aguja Formation of West Texas, and a revision of the Caenagnathidae. Bulletin of the Peabody Museum of Natural History, 54 (1), 23–49.

Lü, J.C. (1999) New material of Ingenia (Barsbold, 1981) from the Nemegt Formation of southwestern Mongolia and its phylogenetic relationships among Oviraptorosauria. Unpublished Master’s Thesis, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing, 50 pp.

Lü, J.C. (2004) Oviraptorid dinosaurs from southern China. Unpublished Ph.D. dissertation, Southern Methodist University, Dallas, 264 pp.

Lü, J.C., Dong, Z.-M., Azuma, Y., Barsbold, R. & Tomida, Y. (2002) Oviraptorosaurs compared to birds. In: Zhou, Z. & Zhang, F. (Eds.), Proceedings of the 5th Symposium of the Society of Avian Paleontology and Evolution. Science Press, Beijing, pp. 175–189.

Lü, J.C., Tomida, Y., Azuma, Y., Dong, Z.M. & Lee, Y.N. (2004) New oviraptorid dinosaur (Dinosauria: Oviraptorosauria) from the Nemegt Formation of southwestern Mongolia. Bulletin of the Natural Science Museum, Series C (Tokyo), 30, 95–130.

Lü, J.C., Tomida, Y., Azuma, Y., Dong, Z.M. & Lee, Y.N. (2005a) Nemegtomaia gen. nov., a new generic name for the oviraptorosaurian dinosaur Nemegtia Lu et al., 2004. Bulletin of the Natural Science Museum, Series C (Tokyo), 31, 51.

Lü, J.C., Huang, D. & Qiu, L.C. (2005b) The pectoral girdle and the forelimb of Heyuannia (Dinosauria: Oviraptorosauria). In: Carpenter, K. (Ed.), The Carnivorous Dinosaurs. Indiana University Press, Bloomington, pp. 256–273.

Lü, J.C., Currie, P.J., Xu, L., Zhang, X., Pu, H. & Jia, S. (2013) Chicken-sized oviraptorid dinosaurs from central China and their ontogenetic implications. Naturwissenschaften, 100(2), 165–175.

Maryańska, T. & Osmólska, H. (1997) The quadrate of oviraptorid dinosaurs. Acta Palaeontologica Polonica, 42, 361–371.

Maryańska, T., Osmólska, H. & Wolsan, M. (2002) Avialan status for Oviraptorosauria. Acta Palaeontologica Polonica, 47, 97–116.

Osborn, H.F. (1924) Three new Theropoda, Protoceratops zone, central Mongolia. American Museum Novitates, 144, 1–12.

Osmólska, H., Currie, P.J. & Barsbold, R. (2004) Oviraptorosauria. In: Weishampel, D.B., Dodson, P. & Osmólska, H. (Eds.), The Dinosauria. 2nd ed. University of California Press, Berkeley, pp. 165–183.

Paul, G.S. (1988) Predatory Dinosaurs of the World: a Complete Illustrated Guide. Simon and Schuster Co., New York, 464 pp.

Qiu, L. & Huang, D. (2001) Dinosaur fossils from the Heyuan Basin in Guangdong Province, China. In: Deng, T. & Wang, Y. (Eds.), Proceedings of the Eighth Annual Meeting of the Chinese Society of Vertebrate Paleontology. China Ocean Press, Beijing, pp. 59–63.

Wang, S., Sun, C., Sullivan, C. & Xu, X. (2013) A new oviraptorid (Dinosauria: Theropoda) from the Upper Cretaceous of southern China. Zootaxa, 3640 (2), 242–257.

Webster, D. (1996) Dinosaurs of the Gobi. National Geographic, 190, 70–89.

Wei, X.F., Pu, H.Y., Xu, L., Lui, D. & Lü, J.C. (2013) A new oviraptorid dinosaur (Theropoda: Oviraptorosauria) from the Late Cretaceous of Jiangxi Province, Southern China. Acta Geologica Sinica (English Edition), 87 (4), 899–904.

Xu, X., Tan, Q.W., Wang, S., Sullivan, C., Hone, D.W.E., Han, F.-L., Ma, Q.-Y., Tan, L. & Xiao, D. (2013) A new oviraptorid from the Upper Cretaceous of Nei Mongol, China, and its stratigraphic implications. Vertebrata PalAsiatica, 51 (2), 85–101.

Zanno, L.E. & Sampson, S.D. (2005) A new oviraptorosaur (Theropoda, Maniraptora) from the Late Cretaceous (Campanian) of Utah. Journal of Vertebrate Paleontology, 25 (4), 897–904.

io.sS�be�� �� (Accessed 10 August 2013)

Rohlf, F.J. (2009) TpsUTIL1.44. Department of Ecology and Evolution, State University of New York at Stony Brook. Available from: (Accessed 10 August 2013)

Skorikov, A.S. (1914) Les formes nouvelles des bourdons (Hymenoptera, Bombidae) VI. Russkoe éntomologicheskoe Obozrênie, 14, 119–129.

Skorikov, A.S. (1923) Palaearctic bumble bees. Part I. General biology (including zoogeography). Izvestiya Severnoi oblastnoi stantsii zashchity rastenii ot vreditelei, 4, 1–160.

Slice, D.E. (2007) Morpheus. Software for morphometric Research. Department of Ecology and Evolution, State University of New York, Stony Brook, New York. Available from: (Accessed 10 August 2013)

Terzo, M., Valterova, I., Urbanova, K. & Rasmont, P. (2003) De la nécessité de redécrire les phéromones sexuelles des mâles de bourdons (Hymenoptera: Apidae, Bombini) publiées avant 1996 pour leur utilisation en analyse phylogénétique. Phytoprotection, 84, 39–49.

Terzo, M., Valterova, I. & Rasmont, P. (2007) Atypical secretions of the male cephalic labial glands in Bumblebees: The case of Bombus (Rhodobombus) mesomelas GERSTAECKER (Hymenoptera, Apidae). Chemistry & Biodiversity, 4 (7), 1466–1471.

Urbanová, K., Valterová, I., Rasmont, P. & Terzo, M. (2002) Znaèkovací feromony: slovení sekretu labiální vlázy samcù Bombus magnus (Marking pheromones of bumblebees: composition of the labial gland secretion of males of Bombus magnus). Chemické listy, 96, 918–919.

Vogt, O. (1911) Studien über das Artproblem. 2. Mitteilung. Über das Variieren der Hummeln. 2. Teil. (Schluss). Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin, 31–74.

Wappler, T., De Meulemeester, T., Aytekin, A.M., Michez, D., Engel, M. (2012) Geometric morphometric analysis of a new Miocene bumble bee from the Randeck Maar of southwestern Germany. Systematic Entomology, 37, 784–792.

Williams, P.H. (1985) A preliminary cladistic investigation of relationships among the bumble bees (Hymenoptera, Apidae). Systematic Entomology, 10, 239–255.

Williams, P.H. (1994) Phylogenetic relationships among bumble bees (Bombus Latr.): a reappraisal of morphological evidence. Systematic Entomology, 19, 327–344.

Williams, P.H. (2007) The distribution of bumblebee colour patterns world-wide: possible significance for thermoregulation, crypsis, and warning mimicry. Biological Journal of the Linnean Society, 92, 97–118.

Williams, P.H., Cameron, S.A., Hines, H.M., Cederberg, B. & Rasmont, P. (2008) A simplified subgeneric classification of the bumblebees (genus Bombus). Apidologie, 39, 46–74.

Williams, P.H., Tang, Y., Yao, J. & Cameron, S.A. (2009) The bumblebees of Sichuan (Hymenoptera: Apidae, Bombini). Systematics and Biodiversity, 7 (2), 101–190.

Zeldtich, M.L., Swiderski, H.D., Sheets, H.D. & Fink, W.L. (2007) IMP. Integrated Morphometrics Package. Available from: (Accessed 10 August 2013)


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