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A revision of the fern genus Dryopteris (Dryopteridaceae) in sub-Saharan Africa



The morphology of the Dryopteris species occurring in sub-Saharan Africa is discussed. This is followed by a revision of the genus in this region, and the Cape Verde Islands in the Gulf of Guinea. Twenty-six species are recognised for the region.


Dryopteris, Dryopteridaceae, Africa, Pteridophytes

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Adanson, M. (1763) Familles des plantes 2: [i], (1)–(24), [i–iii], [1]–640. Vincent, Paris.

Alston, A.H.G. (1956) New African ferns. Boletim da Sociedade Broteriana, sér. 2, 30: 5–27.

Alston, A.H.G. (1959) The ferns and fern-allies of West tropical Africa, being a second edition of the Flora of West tropical Africa: 1–89. London.

Alston, A.H.G. & Schelpe, E.A.C.L.E. (1952) An annotated checklist of the Pteridophyta of southern Africa. Journal of South African Botany 18(4): 153–176.

Baker, J.G. (1867) in Hooker, W.J. & Baker, J.G., Synopsis filicum 7: 257–304. Hardwicke, London.

Baker, J.G. (1874) in Hooker, W.J. & Baker, J.G., Synopsis filicum, edn 2: [i]–xiv, 9–559, pls 1–9. Hardwicke, London.

Baker, J.G. (1875) On the discovery of Phylica arborea Thouars, a tree of Tristan d'Acunha, in Amsterdam Island in the S. Indian Ocean; with an enumeration of the Phanerogams and Vascular Cryptogams of the Island and of St. Paul. Journal of the Linnean Society, Botany 14: 474–480.

Baker, J.G. (1891) A summary of the new ferns which have been discovered or discribed since 1874. Annals of Botany 5: 181–221, 301–332, 455–500.

Barkly, A.M. (1875) A revised list of the ferns of South Africa. Monthly Magazine 10: 193–207.

Barrington, D.S., Paris, C.A. & Ranker, T.A. (1986) Systematic inference from spores and stomate size in the ferns. American Fern Journal 76: 149–159.

Beck, S.L., Dunlop, R.W. & Fossey, A. (2003) Stoma length and frequency as a measure of ploidy level in black wattle, Acacia mearnsii de Wild. Botanical Journal of the Linnean Society 141: 177–181.

Beddome, R.H. (1883) Handbook to the ferns of British India: [i]–xiv, pls 296, 297, [1]–500, pls 1–295, 298–300. Thacker & Spink, Calcutta.

Benl, G. (1991) The Pteridophyta of Bioko (Fernando Po). (Contributions to a flora of the island). Acta Botanica Barcinonensia 40: 1–106.

Bolle, C. (1866) Die Standorte der Farrn auf den canarischen Inseln. III. Zeitschrift der Gesellschaft für Erdkunde zu Berlin 1: 209–238.

Bonaparte, R. (1923) Afrique tropicale plusieurs collecteurs herbier du Prince Bonaparte. Notes Ptéridologiques 14: 199–235.

Bonaparte, R. (1924) Fougères d'Afrique de l'Herbier National des États-Unis, a Washington. Notes Ptéridologiques 15: 7–58.

Bonaparte, R. (1925) Fougères de Madagascar. Supplément. Notes Ptéridologiques 16: 157–198.

Bory de Saint-Vincent, J.B.G.G.M. (1826) Dictionnaire classique d’histoire naturelle 10 [i – iii], [1] – 642. Paris.

Bory de Saint-Vincent, J.B.G.G.M. (1833) Cryptogamie. In C.P. Bélanger, Voyage aux Indes Orientales. Botanique 2,1: 1–80, pls 1–8. Paris.

Bower, F.O. (1923) The ferns (Filicales). Cambridge University Press. Vol. 1: 1–359.

Braun, A. (1841) Bemerkungen über die Flora von Abyssinien. Flora 24,2 (1841): 705–707.

Brause, G.G.W. & Hieronymus, G.H.E.W. (1910) Pteridophyta, in Mildbraed, G.W.J. Wissenschaftliche Ergebnisse der Deutschen Zentral-Afrika-Expedition 1907–1908, Botanik 2: 1–176. Klinkhart & Diermann, Leipzig.

Buchanan, J. (1875) Revised list of the ferns of Natal. Reprinted from the ‘Natal Colonist’, Natal.

Burrows, J.E. (1990) Southern African ferns and fern allies: [i–x], xi–xiii, 2–359, pls 1–56. Frandsen Publishers, Sandton.

Cardoso, J. (1915) Cryptogamias das ilhas de Cabo Verde. Brotéria (Botanica) 13: 89–107.

Chevalier, A. (1935) Les Iles du Cap Vert, géographie, biogéographie, agriculture. Flore de l’Archipel. Revue de Botanique Appliquée et d’Agriculture Tropicale 15(170–171): 1–336.

Ching, R.C. (1938) A revision of the Chinese and Sikkim-Himalayan Dryopteris with reference to some species from neighbouring regions. Bulletin of the Fan Memorial Institute for Biology (Botany) 8: 275–334, plates VI–VII; 8,6: 363–507.

Ching, R.C. (1966) Three new fern genera. Acta Phytotaxonomica Sinica 11(1): 17–29.

Chiovenda, E. (1935) Raccolte botaniche (Embryophyta diploidalia) fatte dai missionari della consolata nel Kenya. Lavori dell'Istituto Botanico, Università di Modena 6(I/XI): 1–161.

Christ, H. (1897) Die Farnkräuter der Erde: [i]–xii, [1]–388. Jena.

Christenhusz, M.J.M., Zhang, X.-C. & Schneider, H. (2011) A linear sequence of extant families and genera of lycophytes and ferns. Phytotaxa 19: 7–54.

Christensen, C. (1905) Index filicum: [1]–384. Hagerup, Copenhagen.

Christensen, C. (1913) Index filicum. Supplémentum 1906–1912: [i]–iv, [1]–131. Hagerup, Copenhagen.

Christensen, C. (1917) Index filicum. Supplementum préliminaire: [i–iii], [1]–60. Hagerup, Copenhagen.

Christensen, C. (1924) Über die Farne des Kenia und Mt. Aberdare, tropisches Ostafrika, in Fries, R.E. & Fries T.C.E., Beiträge zur Kenntnis der Flora des Kenia, Mt. Aberdare und Mt. Elgon. VI. Notizblatt des Botanischen Gartens und Museums zu Berlin-Dahlem 9: 173–189.

Christensen, C. (1932a) Filices, in Perrier de la Bâthie, H., Catalogue des plantes de Madagascar. Pteridophyta: [1]–72. Pitot, Tananarive.

Christensen, C. (1932b) The Pteridophyta of Madagascar. Dansk Botanisk Arkiv 7: [i]–xv, [1]–253, 80 plates.

Daigobo, S. (1972) Taxonomic studies on the fern genus Polystichum in Japan, Ryukyu, and Taiwan. The Science Report of the Tokyo Kyoiku Diagaku Section B 15(224): 57–80.

Dansereau, P. (1961) Études macaronésiennes. I. Géographie des Cryptogames vasculaires. Agronomia Lusitana 23(3): 151–181.

Davies, K.L. (1991) A brief comparative survey of aerophore structure within the Filicopsida. Botanical Journal of the Linnean Society 107: 115–137.

Desvaux, N.A. (1811) Observations sur quelques nouveaux genres de fougéres et sur plusieures espèces nouvelles de la même famille. Magazin für die neuesten Entdeckungen in der gesammten Naturkunde, Gesellschaft Naturforschender Freunde zu Berlin 5: 297–330, figs 4–7.

Desvaux, N.A. (1827) Prodrome de la familles des Fougères. Mémoires de la Société Linnéenne de Paris 6(2): 171–337.

Edwards, D. & Leistner, O.A. (1971) A degree reference system for citing biological records in. South Africa. Mitteilungen aus der Botanischen Staatssammlung München 10: 501–509.

Exell, A.W. (1944) Catalogue of the vascular plants of São Tomé (with Principe and Annobon): Trustees of the British Museum, London. [i–iv] v–xi, [1]–428.

Fée, A.L.A. (1852) Genera filicum. Exposition des genres de las famille des Polypodiacées (Classe des fougères). Cinquième mémoires sur la famille des fougères: 1–387, t. 387. Bailière & Masson, Paris.

Fraser-Jenkins, C.F. (1982) Dryopteris in Spain, Portugal and Macaronesia. Boletim da Sociedade Broteriana, sér. 2, 55: 175–336.

Fraser-Jenkins, C.R. (1986) A classification of the genus Dryopteris (Pteridophyta: Dryopteridaceae). Bulletin of the British Museum (Natural History), Botany 14(3): 183–218.

Fraser-Jenkins, C.R. (1989) A monograph of Dryopteris (Pteridophyta: Dryopteridaceae) in the Indian subcontinent. Bulletin of the British Museum (Natural History), Botany 18(5): 323–477.

Fraser-Jenkins, C.R. (1996) in Widén, C.-J., Fraser-Jenkins, C., Reichstein, T.,Gibby, M. & Sarvela, J. Phloroglucinol derivatives in Dryopteris sect. Fibrillosae and related taxa (Pteridophyta, Dryopteridaceae). Annales Botanici Fennici 33: 69–100.

Fraser-Jenkins, C.R. (2006),in Shaffer-Fehre, M. (vol. ed.), A revised handbook to the Flora of Ceylon 15(2), in: C.R. Fraser-Jenkins & M.J. Zink, Dryopteridaceae: 100–155. Science Publishers, New Hampshire.

Geiger, J.M.O. & Ranker, T.A. (2005) Molecular phylogenetics and historical biogeography of Hawaiian Dryopteris (Dryopteridaceae). Molecular Phylogenetics and Evolution 34: 392–407.

Goldblatt, P. (1974) Biosystematic studies in Papaver section Oxytona. Annals of the Missouri Botanical Garden 61(2): 264–296.

Gray, A. (1848) A manual of the botany of the Northern United States: i–lxxii, [1]–710. Boston & Cambridge.

Gunn, M. & Codd, L.E. (1981) Botanical exploration of southern Africa: [i–vi], vii–xiv, 1–400. Balkema, Cape Town.

Hieronymus, G.H.E.W. (1895) In H.G.A. Engler, Die Pflanzenwelt Ost-Afrikas und der Nachbargebiete 1, C: [i]–iii, [1]–433, [1]–40, i–iii, 45 plates. Reimers, Berlin.

Holttum, R.E. (1974) Thelypteridaceae of Africa and adjacent islands. Journal of South African Botany 40(2): 123–168.

Hooker, W.J. (1862) Species filicum 4(14): 65–160, pls 231–249. Pamplin, London.

Hooker, W.J. (1863) Species filicum 4(15/16): 161–292, pls 250–280. Pamplin, London.

Itô, H. (1935) Filices Japonenses. II. The Botanical Magazine (Tokyo) 49: 432–437.

Itô, H. (1936a) Filices Japonenses. III. The Botanical Magazine (Tokyo) 50: 32–39.

Itô, H. (1936b) Filices Japonenses. IV. The Botanical Magazine (Tokyo) 50: 67–72.

Itô, H. (1936c) Filices Japonenses. V. The Botanical Magazine (Tokyo) 50: 125–128.

Itô, H. (1939) [1938] Polypodiaceae, Dryopteridaceae I, in Nakai, T. & Honda, M. Nova flora japonica 4. Tokyo and Osaka.

Jacobsen, W.B.G. 1978. Some problems of South African Pteridophyta. Journal of South African Botany 44(2): 157–185.

Jacobsen, W.B.G. (1983) The ferns and fern allies of southern Africa. Butterworth, Durban.

Jeanpert, É. (1910) Fougères récoltées par M. Pobéguin au Fouta Djallon (Côte occidentale d'Afrique). Bulletin du Muséum d'Histoire Naturelle 7: 403–404.

Juslén, A., Väre, H. & Wikström, N. (2011) Relationships and evolutionary origins of polyploid Dryopteris (Dryopteridaceae) from Europe inferred using nuclear pgiC and plastid trnL-F sequence data. Taxon 60: 1284–1294.

Keyserling, A.F.M. (1873) Polypodiacea et Cyatheacea herbarii bungeani: [i]–viii, [1]–74. Engelmann, Leipzig.

Kornaś, J. (1979) Distribution and ecology of the pteridophytes in Zambia. Panstwowe Wydawnictwo Naukowe, Warszawa-Kraków.

Kramer, K.U. (1990) Dryopteridaceae, in Kubitzki, K. (ed.), Kramer, K.U. & Green, P.S. (vol. eds.). The families and genera of vascular plants. I. Pteridophytes and Gymnosperms pp. 101–144. Springer-Verlag, Berlin.

Krasser, F. (1900) Filices, Lycopodiaceae et Selaginellaceae, in A. Zahlbrucker, Plantae Pentherianae. Aufzahlung der von Dr. A. Penther und in seinem Auftrage von P. Krook in Sudafrika gesammelten Pflanzen. Annalen des Kaiserlich-Königlichen Naturhistorischen Hofmuseums Wien 15: 3–6.

Kuhn, F.A.M. (1868) Filices africanae: [i], [1]–233. Engelmann, Leipzig.

Kuhn, F.A.M. (1879) Cryptogamae vasculares, in: Ascherson, F.P.A. Von der Decken's, Reisen in Ost-Afrika in 1851–1861, 3,3 Botany: [1]–91, 5 plates. Engelmann, Leipzig.

Kuntze, O. (1891) Revisio generum plantarum 2: [375]–1011. Felix, Leipzig.

Kunze, G. (1836) Plantarum acotyledonearum Africae australis recensio nova. Linnaea 10: 480–570.

Kunze, G. (1844) Filicum in Promontorio Bonae Spei et ad portum Natalensem a Gueinzio nuperius collectarum. Linnaea 18: 113–124.

Lehtonen, S. 2011. Towards resolving the complete fern tree of life. Plos ONE 6,10, e24851. doi:10.1371/

Li, C.-X. & Lu, S.-G. (2006) Phylogenetics of Chinese Dryopteris (Dryopteridaceae) based on the chloroplast rps4-trnS sequence data. Journal of Plant Research 119: 589–598.

Lin, B.-L. & De Vol, C.E. (1977) The use of stipe characters in fern taxonomy I. Taiwania 22: 91–99.

Lin, B.-L. & De Vol, C.E. (1978) The use of stipe characters in fern taxonomy II. Taiwania 23: 77–95.

Liu, H.-M., Zhang, X.-C., Wang, W., Qiu, Y.-L. & Chen, Z.-D. (2007) Molecular phylogeny of the fern family Dryopteridaceae inferred from chloroplast rbcL and atpB genes. International Journal of Plant Science 168,9: 1311–1323.

Lobin, W., Fisher, E. & Ormonde, J. (1982) The ferns and fern-allies (Pteridophyta) of the Cape Verde Islands, West Africa. Nova Hedwigia 115: 1–114.

Lobin, W., Fischer, E. & Ormonde, J. (1998) The ferns and fern-allies (Pteridophyta) of the Cape Verde Islands, West Africa. Nova Hedwigia, Beihefte 115: 1–115.

Manton, I. (1959) Cytological Information on the ferns of west tropical Africa, in Alston, A.H.G. The ferns and fern-allies of west tropical Africa, being a second edition of the Flora of West Tropical Africa: 75–81. London.

McKen, M.J. (1869) The ferns of Natal. P. Davis & Sons, Pietermaritzburg.

McNeill, J., Barrie, F.R., Burdet, H.M., Demolin, V., Hawksworth, D.L., Marhold, K., Nicholson, D.H., Prado, J., Silva, P.C., Skog, J.E., Wiersema, J.H. & Turland, N.J. (2006) International Code of Botanical Nomenclature (Vienna Code) adopted by the Seventeeth International Botanical Congress, Vienna, Austria, July 2005. Regnum Vegetabile 146: i–xviii, 1–568. Koeltz Scientific Books, Königstein.

Metcalfe, C.R. & Chalk, L. (1950) Anatomy of the dicotyledons. Clarendon Press, Oxford. 2 vols., 1500 p.

Michaux, A. (1803) Flora boreali-americana 2: [i, iii], [1]–340, pls 30–51. Paris.

Milde, J. (1867) Filices Europae et Atlantidis, Asiae Minoris et Siberiae: [i]–iv, [1]–311. Felix, Leipzig.

Moore, T. (1853a) List of Mr. Plant's Natal ferns. Journal of Botany (W.J. Hooker, ed.) 5: 225–229.

Moore, T. (1853b) The Uncomo-como, or I'komo-komo, of Natal. Journal of Botany (W.J. Hooker, ed.) 5: 311.

Moore, T. (1855) New garden ferns – No. II. The Gardeners Chronicle & Agricultural Gazette 1855: 468–469.

Moore, T. (1857) Index filicum 1: xcv, xcvi, pl. LXXVIIB. Pamplin, London.

Mucina, L., Hoare, D.B., Lötter, M.C., du Preez, P.J., Rutherford, M.C., Scott-Shaw, C.R., Bredenkamp, G.J., Powrie, L.W., Scott, L., Camp, K.G.T., Cilliers, S.S., Bezuidenhout, H., Mostert, T.H., Siebert, S.J., Winter, P.J., Burrows, J.E., Dobson, L., Ward, R.A., Stalmans, M., Oliver, E.G.H., Siebert, F., Schmidt, E., Kobesi, K., Kose, L. (2006) Grassland Biome, in Mucina, L. & Rutherford, M.C. (eds.). The vegetation of South Africa, Lesotho and Swaziland. Strelitzia 19: 349–436. South African National Biodiversity Institute, Pretoria.

Newman, E. (1851) Proposed addition of three new species and three new genera of British ferns. The Phytologist 4: 368–371, appendix i–xxiii.

Page, C.N. (1968) Cytotaxonomic and anatomical studies on certain Pteridophyta. Ph.D. Thesis, Universiy of Newcastle (unpublished).

Pappe, C.W.L. & Rawson, R.W. (1858) Synopsis filicum Africae australis: [i]–viii, [1]–57. Brittain, Cape Town.

Peter, A. (1929) Flora von Deutsch-Ostafrika. Feddes Repertorium, Beiheft 40(1): 1–98, Anhang: 1–9, t. I–V.

Pichi Sermolli, R.E.G. (1951) On Desvaux’s Aspidium oliganthum and Aspidium oligodonton. Webbia 8: 147–154.

Pichi Sermolli, R.E.G. (1977) Fragmenta pteridographiae VII. Webbia 32(1): 69–93.

Pichi Sermolli, R.E.G. (1984) A new species of Dryopteris from tropical East Africa. Webbia 37(2): 329–339.

Pichi Sermolli, R.E.G. (1985) A contribution to the knowledge of the Pteridophyta of Rwanda, Burundi, and Kivu (Zaïre)—II. Bulletin du Jardin Botanique National de Belgique 55(1/2): 123–206.

Presl, C. (1836) Tentamen pteridographiae: [i], [1]–290, pls 1–12. Prague.

Presl, C. (1851) Epimeliae botanicae. Abhandlungen der Königlichen Böhmischen Gesellschaft der Wissenschaften, ser. 5, 6: 361–624, t. 1–15. Reprinted as Epimeliae botanicae: 1–264, t. 1–15 (1851). Haase, Prague.

Richard, L.C.M. (1801) In F. Marthe, Catalogue des plantes du jardin médicinale de Paris: [1]–144. Gabon, Paris.

Rosenstock, E. (1907) Filices novae. Feddes Repertorium 4: 2–6.

Roth, A.W. (1799) Tentamen florae germanicae 3,1: [i], [1]–102. Leipzig.

Roux, J.P. (1986) A review and typification of some of Kunze’s newly described South African Pteridophyta published in his Acotyledonearum Africae Australis Recencio Nova. Botanical Journal of the Linnean Society 92(4): 343–381.

Roux, J.P. (2000) The genus Polystichum (Dryopteridaceae) in Africa. Bulletin of the natural History Museum London (Botany) 30: 33–79.

Roux, J.P. (2001) Conspectus of southern African Pteridophyta. Southern African Botanical Diversity Network Report 13: 1–223. SABONET, Pretoria.

Roux, J.P. (2002) A new species of Dryopteris (Dryopteridaceae: Pteropsida) from East Africa. Kew Bulletin 57(3): 735–739.

Roux, J.P. (2003) A new Dryopteris (Pteropsida: Dryopteridaceae) species from south-central Africa. Botanical Journal of the Linnean Society 142(2): 237–242.

Roux, J.P. (2004a) Dryopteris filipaleata (Pteropsida: Dryopteridaceae), a new species from tropical East Africa. Bothalia 34(1): 27–30.

Roux, J.P. (2004b) Three new species of Dryopteris (Pteropsida: Dryopteridaceae) from West Africa. Kew Bulletin 59(2): 207–217.

Roux, J.P. (2004c) A re-evaluation of the Dryopteris fadenii (Pteropsida: Dryopteridaceae) complex, with the new species D. rodolfii. Webbia 59(1): 133–147.

Roux, J.P. (2004d). Dryopteris gorgonea (Pteropsida: Dryopteridaceae), a new species from the Cape Verde Islands. Bothalia 34(1): 32–35.

Roux, J.P. (2005) [2004]. New species in the fern genus Dryopteris (Dryopteridaceae) from the islands in the Gulf of Guinea (West Africa). Systematics and Biodiversity 2(2): 163–168.

Roux, J.P. 2009. Synopsis of the Lycopodiophyta and Pteridophyta of Africa, Madagascar, and neighbouring islands. Strelitzia 32: 296. South African National Biodiversity Institute, Pretoria.

Roux, J.P. (2011) The fern genera Dryopteris and Nothoperanema (Dryopteridaceae) in Madagascar and neighbouring Indian Ocean islands, including Saint Paul. Adansonia, sér. 3, 33(1): 7—67.

Schelpe, E.A.C.L.E. (1967) New taxa of Pteridophyta from south eastern tropical Africa. Boletim da Sociedade Broteriana, sér. 2, 41: 203–217.

Schelpe, E.A.C.L.E. (1969) A revised check-list of the Pteridophyta of Southern Africa. Journal of South African Botany 35: 127–140.

Schelpe, E.A.C.L.E. (1970) Pteridophyta, in Exell, A.W. & Launert, E. Flora zambesiaca: [1–5], 6–254. Crown Agents for Overseas Governments and Administrations, London.

Schelpe, E.A.C.L.E. & Anthony, N.C. (1982) New species and new combinations in African Pteridophyta and Orchidaceae, in Anthony, N.C. & Schelpe, E.A.C.L.E. in Contributions from the Bolus Herbarium 10: 143–161.

Schelpe, E.A.C.L.E. & Anthony, N.C. (1986) Pteridophyta, in Leistner, O.A. Flora of southern Africa: [i–iv], v–xv, 1–292. Department of Agriculture and Water Supply, Pretoria.

Schlechtendal, D.F.L. von (1825) Adumbrationes plantarum 2: 17–24, pls 7–12. Berlin.

Schott, H.W. (1834) Genera filicum 2: pls 6–10. Wallishausser, Vienna.

Schrader, H.A. (1824) Illustratio filicum a serenissimo principe neowidensi in Brasilia observatarum, praemissis animadversionibus de hajus familiae structura et oeconomia. Göttingische Gelehrte Anzeigen 1824: 857–872.

Schuettpelz, E. & Pryer, K.M. (2007) Fern phylogeny inferred from 400 leptosporangiate species and three plastid genes. Taxon 56(4): 1037–1050.

Sessa, E.B., Zimmer, E.A. & Givnish, T.J. (2012) Phylogeny, divergence times, and historical biogeography of the New World Dryopteris (Dryopteridaceae). American Journal of Botany 99(4): 730–750.

Sim, T.R. (1891) Handbook of the ferns of Kaffraria: [i]–vi, [vii], [1]–62, [63], pls 1–66. Taylor & Henderson, Aberdeen.

Sim, T.R. (1892) The ferns of South Africa: [i]–iv, [1]–275, pls I–CLIX. Juta, Cape Town.

Sim, T.R. (1915) The ferns of South Africa, edn 2: [i]–ix, [xi], [1]–384, pls 1–186. Cambridge University Press, Cambridge.

Smith, A.R. & Fraser-Jenkins, C.R. (1982) Dryopteris paleacea is a synonym of D. wallichiana. Taxon 31(2): 326–329.

Smith, J.E. (1804) Flora brittanica 3 [i], [915] – 1407, [1–2, index]. Davis, London.

Stolze, R.G. (1981) Ferns and fern allies of Guatemala. Part II. Polypodiaceae. Fieldiana, Botany, N.S. 6: 1–522.

Swartz, O. (1801) Genera et species filicum ordine systematico redactarum. Journal für die Botanik (H.A. Schrader, ed.) 1800(2): 1–136, t. 1, 2.

Tagawa, M. (1938) Nothoperanema Tagawa, a new subgen. of Dryopteris. Acta Phytotaxonomica et Geobotanica 7: 198–200.

Tardieu-Blot, M.-L. (1943) Fougères de Guinée. Bulletin de la Société Botanique de France 90(1/6): 94–97.

Tardieu-Blot, M.-L. (1946). Sur la flore ptéridologique des iles atlantiques. Mémoires de la Société de Biogéographie 8: 325–347.

Tardieu-Blot, M.-L. 1948. Fougères nouvelles d'Afrique récoltées par Mr Schnell. Notulae Systematicae (Paris) 13(4): 370–373.

Tardieu-Blot, M.-L. (1958) 5e Famille.—Polypodiacées (sensu lato) [5(1) Dennstaedtiacées—(10) Aspidiacées], in Humbert, H. (ed.), Flore de Madagascar et des Comores (Plantes Vasculaires) 1: 1–391. Firmin-Didot, Paris.

Tardieu-Blot, M.-L. (1965) A propos de quelques combinaisons et espèces nouvelles de fougères Africaines ou Malgaches. Adansonia 5(4): 493–502.

Tardieu-Blot, M.-L. & Des Abbayes, H. (1951). Contribution a la flore des pteridophytes d’A.O.F. (Guinee et Côte-d-Ivoire) III. Bulletin de l’Institut Française d’Afrique Noire. Série A, sciences naturelles: 372–374.

Tryon, A.F. & Lugardon, B.L. (1990) Spores of the Pteridophyta. Springer-Verlag, New York.

Tryon, R.M. & Tryon, A.F. (1982) Ferns and allied plants with special reference to tropical America. Springer-Verlag, New York.

Viane, R.L.L. (1986) Taxonomic significance of the leaf indument in Dryopteris (Pteridophyta): I. Some North American, Macaronesian and European taxa. Plant Systematics and Evolution 153: 77–105.

Wagner, W.H. & Johnson, D.M. (1983) Trophopod, a common overlooked storage structure of potential systematic value in ferns. Taxon 32(2): 268–269.

Wallich, N. (1829) A numerical list of dried specimens: numbers 1–2153. London.

Watt, J.M. & Breyer-Brandwijk, M.G. 1962. The medicinal and poisonous plants of southern and eastern Africa. Livingstone, Edinburgh.

Webb, P.B. (1849) Spicilegia gorgonea; or a catalogue of all plants as yet discovered in the Cape de Verd Islands from the collections of J.D. Hooker, T. Vogel and other travellers. In W.J. Hooker, Niger Flora: [i–ii], [1]–587. Hippolyte Baillière, London.

White, F. (1983) The vegetation of Africa. UNESCO, Paris.

Widén, C.-J., Faden, R.B., Lounasmaa, M., Vida, G., Von Euw, J. & Reichstein, T. (1973) Die phloroglucide von neun Dryopteris-Arten aus Kenya sowie der D. oligodonta (Desv.) Pic.-Serm. und D. “dilatata” von den Canarischen Inseln. Helvetica Chimica Acta 56(7): 2125–2151.

Widén, C.-J., Fraser-Jenkins, C., Reichstein, T., Gibby, M. & Sarvela, J. (1996) Phloroglucinol derivatives in Dryopteris sect. Fibrillosae and related taxa (Pteridophyta, Dryopteridaceae). Annales Botanici Fennici 33: 69–100.

Widén, C.-J., Fraser-Jenkins, C.R., Reichstein, T. & Sarvela, J. (1999) A survey of phenolic compounds in Dryopteris and related fern genera. Part II. Phloroglucinol derivatives in subgenus Dryopteris (Pteridophyta, Dryopteridaceae). Acta Botanica Fennica 164: 16–54.

Willdenow, C.L. von (1810) Cryptogamia, in Species plantarum, ed. 4, 5(1): 1–542. Berlin.

Wu, S. (2000) Dryopteridaceae. In Flora Reipublicae Polularis Sinicae 5, 1: 1–241. Science Press, Beijing.


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