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New species of Pseudosmittia Edwards, 1932 and new records of Allocladius Kieffer, 1913 (Diptera: Chironomidae, Orthocladiinae) from South America



Three new species of Pseudosmittia Edwards from the Neotropical region, P. sætheri, P. larga and P. trapezoidea, are described and figured as males. Allocladius fortispinatus (Edwards), A. globosus Andersen et al. and A. quadrus Andersen et al. are redescribed on the basis of new material and their geographical distribution updated. A phylogenetic analysis was performed in order to assess the phylogenetic relationships of the new species. As a result, the species P. sætheri and P. larga are found to be sister species belonging to the angusta group. The species P. trapezoidea is the sister group of the clade ((P. propetropis (P. tropisP. lamellata)) and belongs to the brevifurcata group sensu lato.


Pseudosmittia, Allocladius, Orthocladiinae, phylogeny, Sierra de la Ventana, Patagonia, Andes, Argentina, Neotropical region

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Andersen, T., Sæther, O.A. & Mendes, H.F. (2010) Neotropical Allocladius Kieffer, 1913 and Pseudosmittia Edwards, 1932 (Diptera: Chironomidae). Zootaxa, 2472, 1–77.

Ferrington, L.C. & Sæther, O.A. (2011) A revision of the genera Pseudosmittia Edwards, 1932, Allocladius Kieffer, 1913, and Hydrosmittia gen. n. (Diptera: Chironomidae, Orthocladiinae). Zootaxa, 2849, 1–314.

Frenguelli, J. (1950) Rasgos generales de la morfología y la geología de la provincia de Buenos Aires. La Plata, Argentina: Laboratorio de Ensayo de Materiales e Investigaciones Tecnológicas, Serie II, 33, 1–33.

Goloboff, P.A. (1999) Analyzing large data sets in reasonable times: solutions for composite optima. Cladistics, 15, 415–428.

Goloboff, P.A. (2002) Techniques for analyzing large data sets. In: DeSalle, R., Giribet, G.. & Wheeler, W. (eds), Techniques in Molecular Systematics and Evolution. Birkhäuser Verlag, Basel, pp. 70–79.

Goloboff, P.A., Farris, J., Källersjö, M., Oxelmann, B., Ramirez, M. & Szumik, C. (2003) Improvements to resampling measures of group support. Cladistics, 19, 324–332.

Goloboff, P.A., Farris, J.S. & Nixon, K.C. (2008) TNT, a free program for phylogenetic analysis. Cladistics, 24, 1–13.

Hurtado, M.A., Jiménez, J.E. & Cabral, M.G. (2006) Análisis ambiental del partido de La Plata. Aportes al ordenamiento territorial. Instituto de Geomorfología y Suelos CISAUA, Ministerio de Asuntos Agrarios Provincia de Buenos Aires y Facultad de Ciencias Naturales y Museo, La Plata. 134 pp.

Paggi, A.C. (1993) Redescription of Pseudosmittia bilobulata (Edw.) n. comb. (= Spaniotoma (Smittia) bilobulata Edwards 1931) and description of P. neobilobulata sp.n. (Diptera: Chironomidae). Annales de Limnologie, 29, 171–174.

Prevosti, F.J. & Chemisquy, M.A. (2010) The impact of missing data on real morphological phylogenies: influence of the number and distribution of missing entries. Cladistics, 26, 326–339.

Ringuelet, R.A. (1961) Rasgos fundamentales de la zoogeografía de la Argentina. Physis, 22, 151–170.

Sæther, O.A. (1980) Glossary of chironomid morphology terminology (Diptera: Chironomidae). Entomologica scandinavica, Supplement 14, 1–51.

Sæther, O.A. (1990) A review of the genus Limnophyes Eaton from the Holarctic and Afrotropical regions (Diptera: Chironomidae, Orthocladiinae). Entomologica scandinavica, Supplement 35, 1–139.

Siri, A. & Donato, M. (2012) Two new species of Podonomus (Chironomidae: Podonominae) from the decarthrus group from Ventania system, Argentina. Zootaxa, 3548, 39–54.

Spies, M. & Reiss, F. (1996) Catalog and bibliography of Neotropical and Mexican Chironomidae (Insecta, Diptera). Spixiana, Supplement, 22, 61–119.

Strenzke, K. (1950) Systematik, Morphologie und Ökologie der terrestrischen Chironomiden. Archiv für Hydrobiologie, Supplement, 18, 207–414.

Wiens, J.J. (2006) Missing data and the design of phylogenetic analyses. Journal of Biomedical Informatics, 39, 34–42.


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