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Order Opiliones Sundevall, 1833. In: Zhang, Z.-Q. (Ed.) Animal Biodiversity: An Outline of Higher-level Classification and Survey of Taxonomic Richness (Addenda 2013)



The taxonomy of harvestmen is progressing at a fast pace. Many of the figures given in the previous outline Kury (2011) have changed in this short space of two years. The total number of valid species grows slowly because the new synonymies from revisions almost cancel out the new descriptions. The total of 6484 extant species of Opiliones (Kury 2011) has increased to 6534, or less than 1%.



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Alegre-B., A. & L.F. Armas (2012) A new species of Heterolacurbs (Opiliones: Biantidae: Stenostygninae) from Puerto Rico. The Journal of Arachnology, 40, 291–295.

Benavides, L.R. & Giribet, G. (2013) A Revision of Selected Clades of Neotropical Mite Harvestmen (Arachnida, Opiliones, Cyphophthalmi, Neogoveidae) with the Description of Eight New Species. Bulletin of the Museum of Comparative Zoology, 161(1), 1–44.

Boyer, S.L., Clouse, R., Benavides, L.R., Sharma, P., Schwendinger, P.J., Kuranarathna, I. & Giribet, G. (2007) Biogeography of the world: a case study from cyphophthalmid Opiliones, a globally distributed group of arachnids. Journal of Biogeography, 34, 2070–2085.

Bragagnolo, C. & Pinto-da-Rocha, R. (2012) Systematic review of Promitobates Roewer, 1913 and cladistic analysis of Mitobatinae Simon, 1879 (Arachnida: Opiliones: Gonyleptidae). Zootaxa, 3308, 1–48.

Chemeris, A.N. (2012) Three new species of the genus Phalangium Linnaeus, 1758 (Arachnida: Opiliones: Phalangiidae) from Armenia and Uzbekistan. Arthropoda selecta, 21(1), 13– 24.

Chemeris, A.N. & Kovblyuk, M.M. (2012) A new species of the genus Giljarovia Kratochvil, 1959 (Arachnida: Opiliones: Nemastomatidae) from the Crimea. Arthropoda Selecta, 21(3), 243–246.

Clouse, R.M. (2012) The lineages of Stylocellidae (Arachnida: Opiliones: Cyphophthalmi). Zootaxa, 3595, 1–34.

Clouse, RM, General, DM, Diesmos, AC & Giribet, G (2011) An old lineage of Cyphophthalmi (Opiliones) discovered on Mindanao highlights the need for biogeographical research in the Philippines. Journal of Arachnology, 39(1), 147–153.

Clouse, R.M. & Giribet, G. (2012) On the Cyphophthalmi (Arachnida, Opiliones) types from the Museo Civico di Storia Naturale ‘Giacomo Doria’. Bulletin of the Museum of Comparative Zoology, 160(5), 241–257.

Clouse, R.M. & Schwendinger, P.J. (2012) Leptopsalis foveolata sp n., a new species of Stylocellidae from Thailand that displays a novel morphological feature in the suborder Cyphophthalmi (Arachnida, Opiliones). Revue Suisse de Zoologie, 119(4) 529–546.

Crawford, R.L. (1992) Catalogue of the genera and type species of the harvestman superfamily Phalangioidea (Arachnida). Burke Museum Contributions in Anthropology and Natural History, 8, 1–60.

Cruz-López, J.A. & Francke, O.F. (2012) Una nueva especie del género Paramitraceras Pickard-Cambridge (Opiliones: Laniatores: Stygnopsidae) de Veracruz, México. Revista Ibérica de Aracnología, 20, 17–23.

Cruz-López, J.A. & Francke, O.F. (2013) Two new species of the genus Paramitraceras Pickard-Cambridge, 1905 (Opiliones: Laniatores: Stygnopsidae) from Chiapas, Mexico. Zootaxa, 3641 (4), 481–490.

DaSilva, M.B. & Gnaspini, P. (2010) A systematic revision of Goniosomatinae (Arachnida : Opiliones : Gonyleptidae), with a cladistic analysis and biogeographical notes. Invertebrate Systematics, 23(6) (“2009”), 530–624.

DaSilva, M.B. & Pinto-da-Rocha, R. (2012) Descriptions of Thereza murutinga sp. nov. and Pristocnemis caipira sp. nov., and new records of Caelopyginae (Opiliones: Laniatores: Gonyleptidae). Zootaxa, 3317: 25–38.

Dunlop, J.A. (2007) Paleontology. In: Pinto-da-Rocha, R., Machado, G. & Giribet, G. (Eds.), Harvestmen: the biology of the Opiliones. x + 597 pages. Harvard University Press, Cambridge and London, pp 247–265.

Dunlop, J.A., Anderson, L.I. Kerp, H. & Hass, H. (2004) A harvestman (Arachnida: Opiliones) from the Early Devonian Rhynie cherts, Aberdeenshire, Scotland. Transactions of the Royal Society of Edinburgh, Earth science, 94, 341–354.

Dunlop, J.A. & L.I. Anderson (2005) A fossil harvestman (Arachnida, Opiliones) from the Misissippian of East Kirkton, Scotland. The Journal of Arachnology, 33, 482–489.

Dunlop, J.A. Bartel C. & Mitov P.G. (2012) An enigmatic spiny harvestman from Baltic amber. Fossil Record, 15(2), 91–101.

Garwood, R.J., Dunlop, J.A., Giribet, G. & Sutton, M.D. (2011) Anatomically modern Carboniferous harvestmen demonstrate early cladogenesis and stasis in Opiliones. Nature Communications, 2, 444.

Giribet, G. (2000) Catalogue of the Cyphophthalmi of the World (Arachnida, Opiliones). Revista Ibérica de Aracnología, 2, 49–76.

Giribet, G. & Kury, A.B. (2007) Chapter 3. Phylogeny and Biogeography. In: Pinto-da-Rocha, R., Machado, G. & Giribet, G. (Eds.), Harvestmen: the biology of the Opiliones. x + 597 pages. Harvard University Press, Cambridge and London, pp. 62–87.

Giribet, G., Vogt, L., Pérez, A., Sharma, P. & Kury, A. (2010) A multilocus approach to harvestmen phylogeny with emphasis on biogeography and the phylogeny of Laniatores. Cladistics, 26(4), 408–437.

Giribet, G., Sharma, P. P., Benavides, L. R., Boyer, S. L., Clouse, R. M., de Bivort, B. L., Dimitrov, D., Kawauchi, G.Y., Murienne, J.Y. & Schwendinger, P.J. (2012). Evolutionary and biogeographical history of an ancient and global group of arachnids (Arachnida: Opiliones: Cyphophthalmi) with a new taxonomic arrangement. Biological Journal of the Linnean Society, 105, 92–130.

González-Luque, C. & Labrada, L. (2012) A new cave-dwelling endemic Ischyropsalis C.L. Koch, 1839 (Opiliones: Dyspnoi: Ischyropsalididae) from the karstic region of Cantabria (Spain). Zootaxa, 3506, 26–42.

Huang, D.-Y., Selden, P.A. & Dunlop, J.A. (2009) Harvestmen (Arachnida: Opiliones) from the Middle Jurassic of China. Naturwissenschaften, 96, 955–962.

Karaman, I.M. (2012) Revision of the genus Karripurcellia Giribet 2003 (Opiliones: Cyphophthalmi: Pettalidae). Biologia Serbica, 34(1–2), 89–99.

Karaman, I.M. (2013) Nemaspela ladae sp. n., a new troglobitic nemastomatid (Opiliones, Dyspnoi, Nemastomatidae) from a Dinaric cave. Zootaxa, 3694(3), 240–248.

Kury, A.B. (2003) Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Ibérica de Aracnología, vol. especial monográfico, 1, 1–337.

Kury, A.B. (2011) Order Opiliones Sundevall, 1833. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa, 3148, 112–114.

Kury, A.B. (2012a) A new genus of Cranaidae from Ecuador (Opiliones: Laniatores). Zootaxa 3314, 31–44.

Kury, A.B. (2012b) First report of the male of Zamora granulata Roewer 1928, with implications on the higher taxonomy of the Zamorinae (Opiliones, Laniatores, Cranaidae). Zootaxa, 3546: 29–42.

Kury, A.B. (2013) The first species of Roquettea from Maranhão, Brazil (Opiliones, Cosmetidae, Discosomaticinae). Zoologia, In press.

Kury, A.B. & Ferreira, C.P. (2012) Two new species of Roquettea Mello-Leitão, 1931 from northern Brazil (Opiliones: Laniatores: Cosmetidae), Zootaxa, 3328, 35–46.

Mendes, A.C. (2009) Avaliação do status sistemático dos táxons supragenéricos da infra-ordem Insidiatores Loman, 1902 (Arachnida, Opiliones, Laniatores). Unpublished Ph.D. thesis. Museu Nacional/UFRJ, Programa de Pós-Graduação em Zoologia, Rio de Janeiro, xvii + 108 p.

Mendes, A.C. (2011) Phylogeny and taxonomic revision of Heteropachylinae (Opiliones: Laniatores: Gonyleptidae). Zoological Journal of the Linnean Society, 163, 437–483.

Mendes, A.C. & Barros, C.M.L. (2013) Description and phylogenetic position of a new species of Metarthrodes (Opiliones: Gonyleptidae: Caelopyginae) from Bahia, northeastern Brazil. Zoologia, 30 (3), 317–323.

Mendes, A.C. & Kury, A.B. (2012) Notes on the systematics of the Triaenonychinae from Madagascar with description of a new species of Acumontia Loman (Opiliones: Laniatores). Zootaxa, 3593, 40–58.

Ochoa, J.A. & Pinto-da-Rocha, R. (2013) Three new Peruvian species of Protimesius (Opiliones: Laniatores: Stygnidae). The Journal of Arachnology, 41,197–204.

Pinto-da-Rocha, R., Benedetti, A.R. Vasconcelos, E.G. & Hara. M.R. (2012) New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys, 198, 25–68.

Pinto-da-Rocha, R. & Tourinho, A.L.M. (2012) Two new genera, ten new species and new records of Amazonian Stygnidae Simon, 1879 (Opiliones: Laniatores). Zootaxa, 3340, 1–28.

Polydoro, N.S. & Pinto-da-Rocha, R. (2012) New species of Fissiphallius Martens 1988 from Brazil and notes on the morphology of Fissiphalliidae (Arachnida, Opiliones). Zootaxa, 3583, 77–82.

Richart, C.H. & Hedin, M. (2013) Three new species in the harvestmen genus Acuclavella (Opiliones, Dyspnoi, Ischyropsalidoidea), including description of male Acuclavella quattuor Shear, 1986. ZooKeys, 311, 19–68.

Schönhofer, A.L. (2013) A taxonomic catalogue of the Dyspnoi Hansen and Sørensen, 1904 (Arachnida: Opiliones). Zootaxa, 3679 (1), 1–68.

Schönhofer A.L. & Martens, J. (2012) The enigmatic Alpine opilionid Saccarella schilleri gen. n., sp. n. (Arachnida: Nemastomatidae)—isolated systematic placement inferred from comparative genital morphology. Organisms Diversity Evolution, 12 (4), 409– 419.

Sharma, P. (2012) New Australasian Zalmoxidae (Opiliones: Laniatores) and a new case of male polymorphism in Opiliones. Zootaxa, 3236, 1–35.

Sharma, P.P., Buenavente, P.A.C., Clouse, R.M., Diesmos, A.C. & Giribet, G. (2012) Forgotten gods: Zalmoxidae of the Philippines and Borneo (Opiliones: Laniatores). Zootaxa, 3280, 29–55.

Sharma, P. & Giribet, G. (2011) The evolutionary and biogeographic history of the armoured harvestmen – Laniatores phylogeny based on ten molecular markers, with the description of two new families of Opiliones (Arachnida). Invertebrate Systematics, 25, 106–142.

Shultz, J.W. (2012) The identity of Hadrobunus grandis: reassignment of Leiobunum aurugineum to H. grandis and H. nonsacculatus new species (Opiliones: Sclerosomatidae: Leiobuninae). Journal of Arachnology, 40(3), 296–303.

Silva, M.P. Hara, M.R. & Pinto-da-Rocha, R. (2013) Revision of the South American Fonckia (Opiliones: Gonyleptidae: Pachylinae) with the description of two new species. Zoologia, 30(2), 227–237.

Snegovaya, N.Y. & Marusik, Y.M. (2012) New species and collections of Opiliones (Arachnida) from Turkey. Acta Arachnologica, 61(2), 59–70.

Taylor, C.K. (2011) Revision of the genus Megalopsalis (Arachnida: Opiliones: Phalangioidea) in Australia and New Zealand and implications for phalangioid classification. Zootaxa, 2773, 1–65.

Villarreal-M., O. & Kury, A.B. (2012) Licornus Roewer, 1932: newly transferred to Ampycinae and first record of the family Gonyleptidae (Opiliones: Laniatores) from Venezuela. Zootaxa, 3544, 71–78.

Zhang, C. & Zhang, F. (2012a) On the subfamilial assignment of Platybunoides (Opiliones: Eupnoi: Phalangiidae), with the description of a new species from China. Zootaxa, 3190, 47–55.

Zhang, C. & Zhang, F. (2012b) A new species of Parabeloniscus (Opiliones: Laniatores: Epedanidae) from China. Zootaxa, 3565, 55–64.

Zhang, C. & Zhang, F. (2013) Description of a new Cladolasma (Opiliones: Nemastomatidae: Ortholasmatinae) species from China. Zootaxa, 3691(4), 443–452.

Zhang, C., Kury, A.B. & Zhang, F. (2013) Notes on Bonea Roewer, 1914 and Lomanius Roewer, 1923 (Opiliones: Laniatores: Podoctidae), with the description of three new species from China. Zootaxa, 3630 (2), 201–224.


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