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The Higher Classification of the Ant Subfamily Ponerinae
(Hymenoptera: Formicidae), with a Review of Ponerine Ecology and Behavior



The tribal and generic classification of the diverse ant subfamily Ponerinae (Hymenoptera: Formicidae) is revised to reflect recent molecular phylogenetic information and a reappraisal of ponerine morphological diversity. The monogeneric tribe Thaumatomyrmecini (Thaumatomyrmex) is newly synonymized under Ponerini (syn. nov.), and the diverse genus Pachycondyla is fragmented into 19 genera, largely along the lines of its junior synonyms: Bothroponera, Brachyponera (gen. rev.), Ectomomyrmex (gen. rev.), Euponera (gen. rev.), Hagensia (gen. rev.), Megaponera (gen. rev.), Mesoponera (gen. rev.), Neoponera (gen. rev.), Ophthalmopone (gen. rev.), Pachycondyla, Paltothyreus (gen. rev.), Pseudoneoponera (gen. rev.), Pseudoponera (gen. rev.), and 6 new genera: Austroponera (gen. nov.), Buniapone (gen. nov.), Fisheropone (gen. nov.), Mayaponera (gen. nov.), Parvaponera (gen. nov.) and Rasopone (gen. nov.). Some junior synonyms of Pachycondyla are transferred to junior synonym status under other genera: Wadeura as a junior synonym of Cryptopone (syn. nov.), and both Termitopone and Syntermitopone as junior synonyms of Neoponera (syn. nov.). A new genus, Iroponera (gen. nov.), based on the new species Iroponera odax (sp. nov.), is described from Australia. Molecular and morphological justifications for these taxonomic changes are given alongside discussions of phylogenetic relationships. Keys to the world genera of Ponerinae are provided, and morphological diagnoses and species lists are given for each genus. Finally, the available information on ponerine ecology and behavior is reviewed and synthesized.



World revision, ants, taxonomy, Ponerinae

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Abe, T. & Uezu, K. (1977) Biology of Diacamma rugosum (Le Guillou) in the Ryukyu Islands with special reference to foraging behaviour. Proceedings of the 8th Congress of the IUSSI, Wageningen, pp. 142–143.

Acosta-Avalos, D., Esquivel, D.M.S., Wajnberg, E., de Barros, H.G.P.L., Oliveira, P.S. & Leal, I. (2001) Seasonal patterns in the orientation system of the migratory ant Pachycondyla marginata. Naturwissenschaften, 88, 343–346.

Acosta-Avalos, D., Wajnberg, E., Oliveira, P.S., Leal, I., Farina, M. & Esquivel, D.M.S. (1999) Isolation of magnetic nanoparticles from Pachycondyla marginata ants. Journal of Experimental Biology, 202, 2687–2692.

Admane, L., Benatchba, K., Koudil, M., Siad, L. & Maziz, S. (2006) AntPart: an algorithm for the unsupervised classification problem using ants. Applied Mathematics and Computation 180, 16–28.

Agarwal, V.M. & Rastogi, N. (2008) Role of floral repellents in the regulation of flower visits of extrafloral nectary-visiting ants in an Indian crop plant. Ecological Entomology, 33, 59–65.

Agbogba, C. (1984) Tandem running behavior in 2 species of ponerine ants - Mesoponera caffraria (Smith) and Hypoponera sp. (Hymenoptera, Formicidae). Insectes Sociaux, 31 (3), 264–276.

Agbogba, C. (1985) Quelques observations sur le comportement de chasse de la ponerine Mesoponera caffraria Smith (Hymenoptera, Formicidae). Actes des Colloques Insectes Sociaux, 2, 235–238.

Agbogba, C. (1990) Comportement des ouvrieres chasseresses de la fourmi Pachycondyla caffraria. Actes des Colloques Insectes Sociaux, 6, 147–150.

Agbogba, C. (1991) Observations on the signals preparing for larval feeding in the ponerine ant Pachycondyla caffraria (Smith). Insectes Sociaux, 38 (4), 439–442.

Agbogba, C. (1992) Settlement in the prey termite nest by the ponerine ant Pachycondyla caffraria (Smith), and tandem running signal analysis for the following ant. Journal of Ethology, 10 (2), 133–137.

Agbogba, C. (1994) Absence of temporal polyethism in the ponerine ant Pachycondyla caffraria (Smith) (Hymenoptera, Formicidae) - early specialization of the foragers. Behavioural Processes, 32 (1), 47–52.

Agbogba, C. & Howse, P.E. (1992) Division of labor between foraging workers of the ponerine ant Pachycondyla caffraria (Smith) (Hymenoptera, Formicidae). Insectes Sociaux, 39 (4), 455–458.

Allard, H.A. (1951) Dinoponera gigantea (Perty), a vicious stinging ant. Journal of the Washington Academy of Sciences, 41, 88–90.

Allard, D., Ito, F., Gobin, B., Tsuji, K. & Billen, J. (2005) Differentiation of the reproductive tract between dominant and subordinate workers in the Japanese queenless ant Diacamma sp. Acta Zoologica, 86, 159–166.

Allard, D., Gobin, B. & Billen, J. (2007) Timing of sperm transfer in Diacamma pallidum. Physiological Entomology, 32, 382–387.

Allard, D., Gobin, B., Ito, F., Tsuji, K. & Billen, J. (2002) Sperm transfer in the Japanese queenless ant Diacamma sp. (Hymenoptera: Formicidae). Netherlands Journal of Zoology, 52 (1), 77–86.

André, E. (1881) Catalogue raisonné des Formicides provenant du voyage en Orient de M. Abeille de Perrin et description des espèces nouvelles. Annales de la Société Entomologique de France, Series 6, 1, 53–78.

André, E. (1890) Matériaux pour servir à la faune myrmécologique de Sierra-Leone (Afrique occidentale). Revue d’Entomologie (Caen), 9, 311–327.

André, E. (1892) Matériaux myrmécologiques. Revue d’Entomologie (Caen), 11, 45–56.

André, J.-P., Peeters, C. & Doums, C. (2001) Serial polygyny and colony genetic structure in the monogynous queenless ant Diacamma cyaneiventre. Behavioral Ecology and Sociobiology, 50, 72–80.

André, J.-B., Peeters, C., Huet, M. & Doums, C. (2006) Estimating the rate of gamergate turnover in the queenless ant Diacamma cyaneiventre using a maximum likelihood model. Insectes Sociaux, 53, 233–240.

Araújo, A. & Rodrigues, Z. (2006) Foraging behavior of the queenless ant Dinoponera quadriceps Santschi (Hymenoptera: Formicidae). Neotropical Entomology, 35 (2), 159–164.

Arnold, G. (1915) A monograph of the Formicidae of South Africa. Part I. Ponerinae; Dorylinae. Annals of the South African Museum, 14, 1–159.

Arnold, G. (1916) A monograph of the Formicidae of South Africa. Part II. Ponerinae, Dorylinae. Annals of the South African Museum, 14, 159–270.

Arnold, G. (1926) A monograph of the Formicidae of South Africa. Appendix. Annals of the South African Museum, 23, 191–295.

Arnold, G. (1946) New species of African Hymenoptera. No. 6. Occasional Papers of the National Museum of Southern Rhodesia, 2, 49–97.

Arnold, G. (1951) The genus Hagensia Forel (Formicidae). Journal of the Entomological Society of Southern Africa, 14, 53–56.

Arnold, G. (1952) New species of African Hymenoptera. No. 10. Occasional Papers of the National Museum of Southern Rhodesia, 2, 460–493.

Ashmead, W.H. (1905) A skeleton of a new arrangement of the families, subfamilies, tribes and genera of the ants, or the superfamily Formicoidea. Canadian Entomologist, 37, 381–384.

Attygalle, A.B., Steghaus-Kovac, S., Ahmad, V.U., Maschwitz, U., Vostrowsky, O. & Bestmann, H.J. (1991) cis-Isogeraniol, a recruitment pheromone of the ant Leptogenys diminuta. Naturwissenschaften, 78, 90–92.

Attygalle, A.B., Vostrowsky, O. & Bestmann, H.J. (1988) (3R,4S)-4-Methyl-3-heptanol, the trail pheromone of the ant Leptogenys diminuta. Naturwissenschaften, 75, 315–317.

Baena, M.L. (1993) Hormigas cazadoras del género Pachycondyla (Hymenoptera; Ponerinae) de la Isla Gorgona y la planicie Pacifica Colombiana. Boletin del Museo de Entomologi’a de la Universidad del Valle, 1 (1), 13–21.

Baratte, S., Cobb, M., Deutsch, J. & Peeters, C. (2005) Morphological variations in the pre-imaginal development of the ponerine ant Diacamma ceylonense. Acta Zoologica, 86, 25–31.

Baratte, S., Cobb, M. & Peeters, C. (2006a) Reproductive conflicts and mutilation in queenless Diacamma ants. Animal Behaviour, 72, 305–311.

Baratte, S., Peeters, C. & Deutsch, J.S. (2006b) Testing homology with morphology, development and gene expression: sex-specific thoracic appendages of the ant Diacamma. Evolution and Development, 8 (5), 433–445.

Baroni Urbani, C. (1975) Contributo alla conoscenza dei generi Belonopelta Mayr e Leiopelta gen. n. (Hymenoptera: Formicidae). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 48, 295–310.

Baroni Urbani, C., Boyan, G.S., Blarer, A., Billen, J. & Musthak Ali, T.M. (1994) A novel mechanism for jumping in the Indian ant Harpegnathos saltator (Jerdon) (Formicidae, Ponerinae). Experientia, 50, 63–71.

Baroni Urbani, C. & de Andrade, M.L. (2003) The ant genus Thaumatomyrmex in Cuba (Hymenoptera: Formicidae) with description of two new species. Bulletin de la Société Entomologique Suisse, 76, 263–277.

Barros, L.A.C., Mariano, C.S.F., Pompolo, S.G. & Delabie, J.H.C. (2009) Hsc-FA and NOR bandings on chromosomes of the giant ant Dinoponera lucida Emery, 1901 (Hymenoptera: Formicidae). Comparative Cytogenetics, 3, 97–102.

Baudry, E., Peeters, C., Brazier, L., Veuille, M. & Doums, C. (2003) Shift in the behaviours regulating monogyny is associated with high genetic differentiation in the queenless ant Diacamma ceylonense. Insectes Sociaux, 50, 390–397.

Beck, J. & Kunz, B.K. (2007) Cooperative self-defence: Matabele ants (Pachycondyla analis) against African driver ants (Dorylus sp.; Hymenoptera: Formicidae). Myrmecological News, 10, 27–28.

Belshaw, R. & Bolton, B. (1994) A survey of the leaf litter ant fauna in Ghana, West Africa. Journal of Hymenoptera Research, 3, 5–16.

Berghoff, S.M., Maschwitz, U. & Linsenmair, K.E. (2003) Influence of the hypogaeic army ant Dorylus (Dichthadia) laevigatus on tropical arthropod communities. Oecologia, 135, 149–157.

Bernard, F. (1950) Notes sur les fourmis de France. II. Peuplement des montagnes méridionales. Annales de la Société Entomologique de France, 115, 1–36.

Bernard, F. (1953) La réserve naturelle intégrale du Mt Nimba. XI. Hyménoptères Formicidae. Mémoires de l'Institut Français d'Afrique Noire, 19, 165–270.

Bernstein, A. (1861) [Untitled. Introduced by: "Von Herrn Dr. Agath. Berstein in Gadok auf Java ist folgendes Schreiben eingegangen:".]. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 11, 7–8.

Billen, J. (2009) Occurrence and structural organization of the exocrine glands in the legs of ants. Arthropod Structure and Development, 38, 2–15.

Billen, J. & Peeters, C. (1991) Fine structure of the gemma gland in the ant Diacamma australe (Hymenoptera, Formicidae). Belgian Journal of Zoology, 121 (2), 203–210.

Billen, J., Stroobants, Z., Wenseleers, T., Hashim, R. & Ito, F. (2013) Diversity and morphology of abdominal glands in workers of the ant genus Myopias (Formicidae, Ponerinae). Arthropod Structure & Development, 42, 165–172.

Bingham, C.T. (1903) The fauna of British India, including Ceylon and Burma. Hymenoptera, Vol. II. Ants and Cuckoo-wasps. Taylor and Francis, London, 506 pp.

Bitsch, J. & Peeters, C. (1991) Moignons alaires et morphologie thoracique chez l’ouvriere de la fourmi Diacamma australe (Fabricius) (Hym. Formicidae Ponerinae). Bulletin de la Société Entomologique de France, 96, 213–221.

Blum, M.S. (1966) Source and specificity of trail pheromones in Termitopone, Monomorium and Huberia, and their relation to those of some other ants. Journal of Entomology Series A - General Entomology, 41, 155–160.

Bocher, A., Doums, C., Millot, L. & Tirard, C. (2008) Reproductive conflicts affect labor and immune defense in the queenless ant Diacamma sp. “nilgiri”. Evolution, 62 (1), 123–134

Bolton, B. (1974) A revision of the ponerine ant genus Plectroctena F. Smith (Hymenoptera: Formicidae). Bulletin of the British Museum (Natural History) (Entomology), 30, 309–338.

Bolton, B. (1975a) A revision of the ant genus Leptogenys Roger (Hymenoptera: Formicidae) in the Ethiopian region with a review of the Malagasy species. Bulletin of the British Museum (Natural History) (Entomology), 31, 235–305.

Bolton, B. (1975b) A revision of the African ponerine ant genus Psalidomyrmex André (Hymenoptera: Formicidae). Bulletin of the British Museum (Natural History) (Entomology), 32, 1–16.

Bolton, B. (1990) The higher classification of the ant subfamily Leptanillinae (Hymenoptera: Formicidae). Systematic Entomology, 15, 267–282.

Bolton, B. (1994) Identification guide to the ant genera of the world. Harvard University Press, Cambridge, Mass., 222 pp.

Bolton, B. (1995) A new general catalogue of the ants of the world. Harvard University Press, Cambridge, Mass., 504 pp.

Bolton, B. (2003) Synopsis and Classification of Formicidae. Memoirs of the American Entomological Institute, 71, 370 pp.

Bolton, B., Alpert, G., Ward, P.S. & Naskrecki, P. (2006) Bolton’s Catalogue of Ants of the World: 1758-2005. Harvard University Press, Cambridge, Mass., CD-ROM.

Bolton, B. & Brown, W.L. Jr. (2002) Loboponera gen. n. and a review of the Afrotropical Plectroctena genus group (Hymenoptera: Formicidae). Bulletin of the British Museum (Natural History) (Entomology), 71, 1–18.

Bolton, B. & Fisher, B.L. (2008a) The Afrotropical ponerine ant genus Asphinctopone Santschi (Hymenoptera: Formicidae). Zootaxa, 1827, 53–61.

Bolton, B. & Fisher, B.L. (2008b) The Afrotropical ponerine ant genus Phrynoponera Wheeler (Hymenoptera: Formicidae). Zootaxa, 1892, 35–52.

Bolton, B. & Fisher, B.L. (2008c) Afrotropical ants of the ponerine genera Centromyrmex Mayr, Promyopias Santschi gen. rev. and Feroponera gen. n., with a revised key to genera of African Ponerinae (Hymenoptera: Formicidae). Zootaxa, 19, 1–37.

Bolton, B. & Fisher, B.L. (2011) Taxonomy of Afrotropical and West Palaearctic ants of the ponerine genus Hypoponera Santschi. Zootaxa, 2843, 1–118.

Bolton, B., Gotwald, W.H. & Leroux, J.-M. (1979) A new West African ant of the genus Plectroctena with ecological notes. Annales de l’Universite d’Abidjan. Serie E, Ecologie, 9 (1976), 371–381.

Bonavita, A. & Lemasne, G. (1970) Food exchange between adults in ants Mesoponera caffraria (Smith, F.). Comptes Rendus Hebdomadaires des Seances de l Academie Des Sciences Serie D, 270 (11), 1511–1514.

Bonavita, A. & Poveda, A. (1970) Mise en evidence d’une division du travail chez une fourmi primitive. Comptes Rendus de l'Académie des Sciences, Paris, 270, 515–518.

Bonavita, A. & Poveda, A. (1972) Preliminary study on organ joining midgut and hindgut of ant larvae (Hymenoptera, Formicidae). Comptes Rendus Hebdomadaires des Seances de l Academie Des Sciences Serie D, 275 (6), 775–778.

Borgmeier, T. (1920) Zur lebensweise von Odontomachus affinis Guérin. Zeitschrift des Deutschen Vereins für Wissenschaft und Kunst in São Paulo, 1, 31–38.

Borgmeier, T. (1950) A fêmea dichthadiiforme e os estádios evolutivos de Simopelta pergandei (Forel), e a descriça~o de S. bicolor, n. sp. (Hym. Formicidae). Revista de Entomologia (Rio de Janeiro), 21, 369–380.

Borgmeier, T. (1959) Myrmecologische Studien. II. Anais da Academia Brasileira de Ciencias, 31, 309–319.

Brady, S.G. (2003) Evolution of the army ant syndrome: The origin and long-term evolutionary stasis of a complex of behavioral and reproductive adaptations. Proceedings of the National Academy of Sciences (USA), 100 (11), 6575–6579.

Brandão, C.R.F. (1983) Sequential ethograms along colony development of Odontomachus affinis Guérin (Hymenoptera, Formicidae, Ponerinae). Insectes Sociaux, 30 (2), 193–203.

Brandão, C.R.F. (1991) Thaumatomyrmex strips millipedes for prey: a novel predatory behaviour in ants, and the first case of sympatry in the genus (Hymenoptera: Formicidae). Insectes Sociaux, 38, 335–344.

Braun, U., Peeters, C. & Hölldobler, B. (1994) The giant nests of the African stink ant Paltothyreus tarsatus (Formicidae, Ponerinae). Biotropica, 26 (3), 308–311.

Brent, C., Peeters, C., Dietmann, V., Crewe, R. & Vargo, E. (2006) Hormonal correlates of reproductive status in the queenless ponerine ant Streblognathus peetersi. Journal of Comparative Physiology, 192 (3), 315–320.

Brown, W.L. Jr. (1950) Morphological, taxonomic, and other notes on ants. Wasmann Journal of Biology, 8, 241–250.

Brown, W.L. Jr. (1952) Contributions toward a reclassification of the Formicidae. I. Tribe Platythyreini (Hymenoptera). Breviora, 6, 1–6.

Brown, W.L. Jr. (1953a) Characters and synonymies among the genera of ants. Part I. Breviora, 11, 1–13.

Brown, W.L. Jr. (1953b) Characters and synonymies among the genera of ants. Part II. Breviora, 18, 1–8.

Brown, W.L. Jr. (1958) A review of the ants of New Zealand. Acta Hymenopterology, 1, 1–50.

Brown, W.L. Jr. (1960) Contributions toward a reclassification of the Formicidae. III. Tribe Amblyoponini (Hymenoptera). Bulletin of the Museum of Comparative Zoology, 122 (4), 145–230.

Brown, W.L. Jr. (1963) Characters and synonymies among the genera of ants. Part III. Some members of the tribe Ponerini (Ponerinae, Formicidae). Breviora, 190, 1–10.

Brown, W.L. Jr. (1973) A comparison of the Hylean and Congo-West African rain forest ant faunas. In: Meggers, B.J., Ayensu, E.S. & Duckworth, W.D. (Eds.), Tropical forest ecosystems in Africa and South America: a comparative review. Smithsonian Institution Press, Washington, D.C., pp. 161–185.

Brown, W.L. Jr. (1974a) Dolioponera genus nov. Pilot Registry of Zoology, Card No. 31.

Brown, W.L. Jr. (1974b) Dolioponera fustigera species nov. Pilot Registry of Zoology, Card No. 32.

Brown, W.L. Jr. (1975) Contributions toward a reclassification of the Formicidae. V. Ponerinae, tribes Platythyreini, Cerapachyini, Cylindromyrmecini, Acanthostichini, and Aenictogitini. Search Agriculture (Ithaca N. Y.), 5 (1), 1–115.

Brown, W.L. Jr. (1976) Contributions toward a reclassification of the Formicidae. Part VI. Ponerinae, tribe Ponerini, subtribe Odontomachiti. Section A. Introduction, subtribal characters. Genus Odontomachus. Studia Entomologica, 19, 67–171.

Brown, W.L. Jr. (1978) Contributions toward a reclassification of the Formicidae. Part VI. Ponerinae, tribe Ponerini, subtribe Odontomachiti. Section B. Genus Anochetus and bibliography. Studia Entomologica, 20, 549–638.

Brown, W.L. Jr. (2000) Diversity of ants. In: Agosti, D., Majer, J., Alonso, E. & Schultz, T.R. (Eds.), Ants: standard methods for measuring and monitoring biodiversity. Smithsonian Institution Press, Washington, D.C., pp. 45–79.

Buschinger, A. & Maschwitz, U. (1984) Defensive behavior and defensive mechanisms in ants. In: Hermann, H.R. (Ed.), Defensive Mechanims in Social Insects. Praeger, New York, pp. 95–150.

Caperucci, D. & Mathias, M.I.C. (2006) Lipids in oocytes of ants Neoponera villosa (Hymenoptera: Ponerinae). Sociobiology, 47 (2), 531–541.

Carlin, N.F. & Gladstein, D.S. (1989) The “bouncer” defense of Odontomachus ruginodis and other odontomachine ants (Hymenoptera: Formicidae). Psyche (Camb.), 96, 1–19.

Carpenter, F.M. (1930) The fossil ants of North America. Bulletin of the Museum of Comparative Zoology, 70, 1–66.

Casnati, G., Ricca, A. & Pavan, M. (1967) Sulla secrezione difensive delle glandole mandibolari di Palttothyreus tarsatus (Fabr.) (Hymenoptera: Formicidae). Chimica e l’Industria (Milan), 49, 57–58.

Chapman, J.W. & Capco, S.R. (1951) Check list of the ants (Hymenoptera: Formicidae) of Asia. Monographs of the Institute of Science and Technology. Manila, 1, 1–327.

Cho, Y.S., Lee, Y., Lee, C., Yoo, B., Park, H. & Moon, H. (2002) Prevalence of Pachycondyla chinensis venom allergy in an ant-infested area in Korea. Journal of Allergy and Clinical Immunology, 110 (1), 54–57.

Clark, J. (1930) New Formicidae, with notes on some little-known species. Proceedings of the Royal Society of Victoria (n.s.), 43, 2–25.

Clark, V.C., Raxworthy, C.J., Rakotomalala, V., Sierwald, P. & Fisher, B.L. (2005) Convergent evolution of chemical defense in poison frogs and arthropod prey between Madagascar and the Neotropics. Proceedings of the National Academy of Sciences (USA), 102 (33), 11617–11622.

Cokendolpher, J.C., Reddell, J.R., Taylor, S.J., Krejca, J.K., Suarez, A.V. & Pekins, C.E. (2009) Further ants (Hymenoptera: Formicidae) from caves of Texas. Texas Memorial Museum Speleological Monographs, 7, 151–168.

Collingwood, C.A., Tigar, B.J. & Agosti, D. (1997) Introduced ants in the United Arab Emirates. Journal of Arid Environments, 37, 505–512.

Collingwood, C.A. & van Harten, A. (2005) Further additions to the ant fauna (Hymenoptera: Formicidae) of Yemen. Zoology in the Middle East, 35, 73–78.

Colombel, P.P. (1968) An ecto-horome eliciting alarm in ants Odontomachus hematodes (Hym Form Poneridae). Comptes Rendus Hebdomadaires des Seances de l Academie des Sciences Serie D, 266 (8), 806–807.

Colombel, P.P. (1970a) Recherches sur la biologie et l’éthologie d’Odontomachus haematodes L. Hym. Formicoïdea Poneridae. Étude des populations dans leur milieu naturel. Insectes Sociaux, 3, 183–198.

Colombel, P.P. (1970b) Recherches sur la biologie et l’éthologie d’Odontomachus haematodes L. Hym. Formicoïdea Poneridae. Biologie des reines. Insectes Sociaux, 3, 199–204.

Colombel, P.P. (1972) Recherches sur la biologie et l’éthologie d’Odontomachus haematodes L. (Hym. Formicoidea, Poneridae) biologie des ouvrieres. Insectes Sociaux, 3, 171–194.

Colombel, P.P. (1974) Artificial breeding of eggs of Odontomachus haematodes L (Hymenoptera, Formicidae, Ponerinae) and trophogenic differentiation of workers and queens. Comptes Rendus Hebdomadaires des Seances de l Academie des Sciences Serie D, 279 (6), 489–491.

Colombel, P.P. (1978) Biologie d’Odontomachus haematodes L. (Hym. Form.) determinisme de la caste femelle. Insectes Sociaux, 25 (2), 141–151.

Corbara, B. & Déjean, A. (2000) Adaptive behavioral flexibility of the ant Pachycondyla analis (Megaponera foetens) (Formicidae: Ponerinae) during prey capture. Sociobiology, 36, 465–483.

Creighton, W.S. (1929) New forms of Odontoponera transversa. Psyche (Camb.), 36, 150–154.

Creighton, W.S. & Tulloch, G.S. (1930) Notes on Euponera gilva (Roger) (Hymenoptera, Formicidae). Psyche (Camb.), 37 (1), 71–79.

Crewe, R.M. & Fletcher, D.J.C. (1974) Ponerine ant secretions: the mandibular gland secretion of Paltothyreus tarsatus Fabr. Journal of the Entomological Society of Southern Africa, 37 (2), 291–298.

Crewe, R.M., Peeters, C. & Villet, M. (1984) Frequency distribution of worker sizes in Megaponera foetens (Fabricius). South African Journal of Zoology, 19, 247–248.

Crewe, R.M. & Ross, F.P. (1975a) Biosynthesis of alkyl sulfides by an ant. Nature, 254 (5499), 448–449.

Crewe, R.M. & Ross, F.P. (1975b) Pheromone biosynthesis - formation of sulfides by ant Paltothyreus tarsatus. Insect Biochemistry, 5 (6), 839–843.

Crosland, M.W.J. (1994) Millipede myrmecophile, Glyphiulus granulatus, found with the primitive ant Harpegnathos venator. Journal of Entomological Science, 29 (1), 141–142.

Crosland, M.W.J. (1995) Nest and colony structure in the primitive ant, Harpegnathos venator (Smith) (Hymenoptera, Formicidae). Pan-Pacific Entomologist, 71 (1), 18–23.

Cuvillier-Hot, V., Cobb, M., Malosse, C. & Peeters, C. (2001) Sex, age and ovarian activity affect cuticular hydrocarbons in Diacamma ceylonense, a queenless ant. Journal of Insect Physiology, 47, 485–493.

Cuvillier-Hot, V., Gadagkar, R., Peeters, C. & Cobb, M. (2002) Regulation of reproduction in a queenless ant: aggression, pheromones and reduction in conflict. Proceedings of the Royal Society B, 269, 1295–1300.

Cuvillier-Hot, V. & Lenoir, A. (2006) Biogenic amine levels, reproduction and social dominance in the queenless ant Streblognathus peetersi. Naturwissenschaften. 93 (3), 149–153.

Cuvillier-Hot, V., Lenoir, A. & Peeters, C. (2004a) Reproductive monopoly enforced by sterile police workers in a queenless ant. Behavioral Ecology, 15 (6), 970–975.

Cuvillier-Hot, V., Lenoir, A., Crewe, R., Malosse, C. & Peeters, C. (2004b) Fertility signalling and reproductive skew in queenless ants. Animal Behaviour, 68, 1209–1219.

Cuvillier-Hot, V., Renault, V. & Peeters, C. (2005) Rapid modification in the olfactory signal of ants following a change in reproductive status. Naturwissenschaften, 92 (2), 73–77.

Dahbi, A. & Jaisson, P. (1995) An analysis of polyethism in the queenless ant Diacamma sp. (Formicidae, Ponerinae). Sociobiology, 26 (1), 69–81.

Dalla Torre, K.W. von (1893) Catalogus Hymenopterorum hucusque descriptorum systematicus et synonymicus. Vol. 7. Formicidae (Heterogyna). W. Engelmann, Leipzig, 289 pp.

Dantas de Araujo, C.Z., Lachaud, J.-P. & Fresneau, D. (1990) Le systéme reproductif chez une ponérine sans reine: Dinoponera quadriceps Santschi. Behavioral Processes, 22, 101–111.

Davidson, D.W. & Fisher, B.F. (1991) Symbiosis of ants with Cecropia as a function of light regime. In: Huxley, C.R. & Cutler, D.F. (Eds.), Ant-plant Interactions. Oxford University Press, Oxford, UK, pp. 289–309.

Davies, S.J., Villet, M.H., Blomefield, T.M. & Crewe, R.M. (1994) Reproduction and division of labour in Leptogenys schwabi Forel (Hymenoptera Formicidae), a polygynous, queenless ponerine ant. Ethology Ecology and Evolution, 6, 507–517.

Dean, W.R.J. (1989) Foraging and forager-recruitment in Ophthalmopone hottentota Emery Hymenoptera Formicidae. Psyche (Camb.), 96 (1–2), 123–130.

De Andrade, M.L. (2004) A new species of Platythyrea from Dominican amber and description of a new extant species from Honduras (Hymenoptera: Formicidae). Revue Suisse de Zoologie, 111 (3), 643–655.

Déjean, A. (1988) Memory effect on predatory behaviour of Odontomachus troglodytes (Formicidae - Ponerinae). Behavior, 107 (3–4), 131–137.

Déjean, A. (1990) Influence de l’environment pre-imaginal et precoce dans le choix du site de nidification de Pachycondyla (=Neoponera) villosa (Fabr.) (Formicidae, Ponerinae). Behavioural Processes, 21, 107–125.

Déjean, A. (1991) The predatory behavior of Pachycondyla soror (Formicidae, Ponerinae). Entomologia Experimentalis et Applicata, 58 (2), 123–135.

Déjean, A. (1997) Distribution of colonies and prey specialization in the ponerine ant genus Leptogenys (Hymenoptera: Formicidae). Sociobiology, 29 (3), 293–300.

Déjean, A. & Bashingwa, E.P. (1985) La prédation chez Odontomachus troglodytes Santschi (Formicidae-Ponerinae). Insectes Sociaux, 32 (1), 23–42.

Déjean, A., Beugnon, G. & Lachaud, J.P. (1993a) Spatial components of foraging behavior in an African ponerine ant Paltothyreus tarsatus. Journal of Insect Behavior, 6 (3), 271–285.

Déjean, A., Bolton, B. & Durand, J.L. (1997) Cubitermes subarquatus termitaries as shelters for soil fauna in African rainforests. Journal of Natural History, 31, 1289–1302.

Déjean, A. & Corbara, B. (1990a) Predatory behavior of a Neotropical arboricolous ant - Pachycondyla villosa (Formicidae, Ponerinae). Sociobiology, 17 (2), 271–286.

Déjean, A. & Corbara, B. (1990b) Glucidic food for the larvae of Pachycondyla villosa (Formicidae, Ponerinae). Biology of Behavior, 15 (3–4), 117–124.

Déjean, A. & Corbara, B. (1998) Study of different foraging paths of the predatory neotropical ponerine ant Pachycondyla (=Neoponera) villosa (Hymenoptera, Formicidae). Sociobiology, 32 (3), 409–426.

Déjean, A., Corbara, B. & Oliva-Rivera, J. (1990) Mise en evidence d’une forme d’apprentissage dans le comportement de capture des proies chez Pachycondyla (=Neoponera) villosa (Formicidae, Ponerinae). Behaviour, 115 (3–4), 175–187.

Déjean, A., Corbara, B., Orivel, J., Snelling, R.R., Delabie, J.H.C. & Belin-Depoux, M. (2000) The importance of ant gardens in the pioneer vegetal formations of French Guiana (Hymenoptera: Formicidae). Sociobiology, 35 (3), 425–439.

Déjean, A., Durand, J.L. & Bolton, B. (1996) Ants inhabiting Cubitermes termitaries in African rain forest. Biotropica, 28, 701–713.

Déjean, A. & Evraerts, C. (1997) Predatory behavior in the genus Leptogenys: a comparative study. Journal of Insect Behavior, 10 (2), 17–191.

Déjean, A. & Fénéron, R. (1996) Polymorphism and oligogyny in the ponerine ant Centromyrmex bequaerti (Formicidae: Ponerinae). Insectes Sociaux, 43, 87–99.

Déjean, A. & Fénéron, R. (1999) Predatory behaviour in the ponerine ant, Centromyrmex bequaerti: a case of termitolesty. Behavioural Processes, 47, 125–133.

Déjean, A. & Lachaud, J.P. (1991) Polyethism in the ponerine ant Odontomachus troglodytes: interaction of age and interindividual variability. Sociobiology, 18 (2), 177–196.

Déjean, A. & Lachaud, J.P. (1994) Ecology and behavior of the seed-eating ponerine ant Brachyponera senaarensis [sic] (Mayr). Insectes Sociaux, 41, 191–210.

Déjean, A., Lachaud, J.P. & Beugnon, G. (1993b) Efficiency in the exploitation of patchy environments by the ponerine ant Paltothyreus tarsatus - an ecological consequence of the flexibility of prey capture behavior. Journal of Ethology, 11 (1), 43–53.

Déjean, A. & Olmsted, I. (1997) Ecological studies on Aechmea bracteata (Swartz) (Bromeliaceae). Journal of Natural History, 31, 1313–1334.

Déjean, A., Olmsted, I. & Camal, J.F. (1992) Interaction between Atta cephalotes and arboreal ants in the Biosphere reserve Sian Ka’an (Quintana Roo, Mexico): efficient protection of the trees (Hymenoptera, Formicidae). Sociobiology, 20 (1), 57–76.

Déjean, A., Schatz, B., Orivel, J., Beugnon, G., Lachaud, J.P. & Corbara, B. (1999) Feeding preferences in African ponerine ants: A cafeteria experiment (Hymenoptera: Formicidae). Sociobiology, 34 (3), 555–568.

Déjean, A. & Suzzoni, J.P. (1991) La predation chez Plectroctena minor (Formicidae: Ponerinae). Actes des Colloques Insectes Sociaux, 7, 137–144.

Déjean, A. & Suzzoni, J.P. (1997) Surface tension strengths in the service of a Ponerine ant: a new kind of nectar transport. Naturwissenschaften, 84, 76–79.

Déjean, A., Suzzoni, J.P. & Schatz, B. (2001) Behavioral adaptations of an African ponerine ant in the capture of millipedes. Behavior, 138, 981–996.

Déjean, A., Suzzoni, J.P., Schatz, B. & Orivel, J. (2002) Territorial aggressiveness and predation: two possible origins of snapping in the ant Plectroctena minor. Comptes Rendus Biologies, 325, 819–825.

De la Mora, A., Pérez-Lachaud, G. & Lachaud, J.P. (2008) Mandible strike: The lethal weapon of Odontomachus opaciventris against small prey. Behavioral Processes, 78, 64–75.

Delabie, J.H.C. (1995) Inquilinismo simultaneo de duas especies de Centromyrmex em cupinzeiros de Syntermes sp. Revista Brasileira de Entomologia, 39, 605–609.

Delabie, J.H.C., Fresneau, D. & Pezon, A. (2000) Notes on the ecology of Thaumatomyrmex spp. (Hymenoptera: Formicidae: Ponerinae) in southeast Bahia, Brazil. Sociobiology, 36, 571–584.

Denis, D., Orivel, J., Hora, R.R., Chameron, S. & Fresneau, D. (2006a) First record of polydomy in a monogynous ponerine ant: a means to allow emigration between Pachycondyla goeldii nests. Journal of Insect Behavior, 19 (3), 279–291.

Denis, D., Blatrix, R. & Fresneau, D. (2006b) How an ant manages to display individual and colonial signals by using the same channel. Journal of Chemical Ecology, 32, 1647–1661.

Denis, D., Chameron, S., Costille, L., Pocheville, A., Châline, N. & Fresneau, D. (2008) Workers agonistic interactions in queenright and queenless nest of a polydomous ant society. Animal Behavior, 75, 791–800.

Denis, D., Pezon, A. & Fresneau, D. (2007) Reproductive allocation in multinest colonies of the ponerine ant Pachycondyla goeldii. Ecological Entomology, 32, 289–295.

D’Ettorre, P. & Heinze, J. (2005) Individual recognition in ant queens. Current Biology, 15, 2170–2174.

D’Ettorre, P., Heinze, J. & Ratnieks, F.L.W. (2004b) Worker policing by egg eating in the ponerine ant Pachycondyla inversa. Proceedings of the Royal Society B, 271, 1427–1434.

D’Ettorre, P., Heinze, J., Schulz, C., Francke, W. & Ayasse, M. (2004b) Does she smell like a queen? Chemoreception of a cuticular hydrocarbon signal in the ant Pachycondyla inversa. Journal of Experimental Biology, 207, 1085–1091.

D’Ettorre, P., Kellner, K., Delabie, J.H.C. & Heinze, J. (2005) Number of queens in founding associations of the ponerine ant Pachycondyla villosa. Insectes Sociaux, 52, 327–332.

D’Ettorre, P., Tofilski, A., Heinze, J. & Ranieks, F.L.W. (2006) Non-transferable signals on ant queen eggs. Naturwissenschaften, 93, 136–140.

Deyrup, M. & Cover, S. (2004) A new species of Odontomachus ant (Hymenoptera: Formicidae) from inland ridges of Florida, with a key to Odontomachus of the United States. Florida Entomologist, 87 (2), 136–144.[0136:ansooa];2

Dib, G., Ferguson, R.K. & Sljivic, V. (1992) Hypersensitivity to Samsun ant. The Lancet, 339, 552–553.

Dietemann, V. & Peeters, C. (2000) Queen influence on the shift from trophic to reproductive eggs laid by workers of the ponerine ant Pachycondyla apicalis. Insectes Sociaux, 47, 223–228.

Djiéto-Lordon, C., Orivel, J. & Déjean, A. (2001a) Consuming large prey on the spot: the case of the arboreal foraging ponerine ant Platythyrea modesta (Hymenoptera, Formicidae). Insectes Sociaux, 48, 324–326.

Djiéto-Lordon, C., Orivel, J. & Déjean, A. (2001b) Predatory behavior of the African ponerine ant Platythyrea modesta (Hymenoptera: Formicidae). Sociobiology, 38 (3A), 303–315.

Dlussky, G.M. & Fedoseeva, E.B. (1988) Origin and early stages of evolution in ants. In: Ponomarenko, A.G. (Ed.), Cretaceous biocenotic crisis and insect evolution. Nauka, Moskva, pp. 70–144.

Don, W. (2008) Ants of New Zealand. Otago University Press, Otago, New Zealand, 239 pp.

Donisthorpe, H. (1937) A new species of Harpegnathos Jerd., with some remarks on the genus, and other known species (Hym. Formicidae). Entomologist's Monthly Magazine, 73, 196–201.

Donisthorpe, H. (1941) Descriptions of new species of ants from New Guinea. Annals and Magazine of Natural History, 11 (7), 129–144.

Donisthorpe, H. (1942) Descriptions of a few ants from the Philippine Islands, and a male of Polyrhachis bihamata Drury from India. Annals and Magazine of Natural History, 11 (9), 64–72.

Donisthorpe, H. (1943a) The ants (Hym., Formicidae) of Waigeu Island, North Dutch New Guinea. Annals and Magazine of Natural History, 11 (10), 433–475.

Donisthorpe, H. (1943b) A list of the type-species of the genera and subgenera of the Formicidae. [part]. Annals and Magazine of Natural History, 11 (10), 617–688.

Donisthorpe, H. (1943c) A list of the type-species of the genera and subgenera of the Formicidae. [concl.]. Annals and Magazine of Natural History, 11 (10), 721–737.

Donisthorpe, H. (1948) Microbolbos testaceus, a new genus and species of ponerine ant. Entomologist, 81, 170–171.

Doums, C. (1999) Characterization of microsatellite loci in the queenless ponerine ant Diacamma cyaneiventre. Molecular Ecology, 8, 1957–1969.

Doums, C., Cabrera, H. & Peeters, C. (2002) Population genetic structure and male-biased dispersal in the queenless ant Diacamma cyaneiventre. Molecular Ecology, 11, 2251–2264.

Dreier, S., van Zweden, J.S. & D’Ettorre, P. (2007) Long-term memory of individual identity in ant queens. Biology Letters, 3, 459–462.

Duelli, P. & Duelli-Klein, R. (1976) Field studies on homing ability in South American ants. Studia Entomologica, 19, 409–420.

Duffield, R.M. & Blum, M.S. (1973) 4-methyl-3-heptanone - identification and function in Neoponera villosa (Hymenoptera: Formicidae). Annals of the Entomological Society of America, 66 (6), 1357.

Duncan, F.D. (1999) The ponerine ant Pachycondyla (=Ophthalmopone) berthoudi Forel carries loads economically. Physiological and Biochemical Zoology, 72 (1), 71–77.

Duncan, F.D. & Crewe, R.M. (1993) A comparison of the energetics of foraging of three species of Leptogenys (Hymenoptera, Formicidae). Physiological Entomology, 18, 372–378.

Duncan, F.D. & Crewe, R.M. (1994a) Field study on the foraging characteristics of a ponerine ant, Hagensia havilandi Forel. Insectes Sociaux, 41, 85–98.

Duncan, F.D. & Crewe, R.M. (1994b) Group hunting in a ponerine ant, Leptogenys nitida Smith. Oecologia, 97, 118–123.

Düssmann, O., Peeters, C. & Hölldobler, B. (1996) Morphology and reproductive behaviour of intercastes in the ponerine ant Pachycondyla obscuricornis. Insectes Sociaux, 43, 421–425.

Eguchi, K., Bui, T.V. & Janssen, R. (2005) Gastropod guests (Prosobranchia: Pupinidae, and Pulmonata: Subulinidae) associated with the ponerine ant Diacamma sculpturatum complex (Insecta: Hymenoptera: Formicidae). Sociobiology, 45 (2), 307–315.

Eguchi, K., Bui, T.V. & Yamane, S. (2004) A preliminary study on foraging distance and nesting sites of ants in indo-Chinese lowland vegetation (Insects, Hymenoptera, Formicidae). Sociobiology, 43 (3), 445–457.

Ehmer, B. & Gronenberg, W. (1997) Proprioceptors and fast antennal reflexes in the ant Odontomachus (Formicidae, Ponerinae). Cell Tissue Research, 290, 153–165.

Ehmer, B. & Hölldobler, B. (1995) Foraging behavior of Odontomachus bauri on Barro Colorado Island, Panama. Pysche, 102, 215–224.

Emery, C. (1877) Saggio di un ordinamento naturale dei Mirmicidei, e considerazioni sulla filogenesi delle formiche. Bollettino della Societa Entomologica Italiana, 9, 67–83.

Emery, C. (1886) Alcune formiche africane. Bollettino della Societa Entomologica Italiana, 18, 355–366.

Emery, C. (1887) Catalogo delle formiche esistenti nelle collezioni del Museo Civico di Genova. Parte terza. Formiche della regione Indo-Malese e dell'Australia (continuazione e fine). [part]. Annali del Museo Civico di Storia Naturale di Genova, 25 [= (2) (5)], 433–448.

Emery, C. (1889) Formiche di Birmania e del Tenasserim raccolte da Leonardo Fea (1885-87). [part]. Annali del Museo Civico di Storia Naturale di Genova, 27 [= (2) (7)], 485–512.

Emery, C. (1890) Studii sulle formiche della fauna neotropica. Bollettino della Societa Entomologica Italiana, 22, 38–80.

Emery, C. (1892) [1891]. Voyage de M. Ch. Alluaud dans le territoire d'Assinie (Afrique occidentale) en juillet et août 1886. Formicides. Annales de la Société Entomologique de France, 60, 553–574.

Emery, C. (1893a) [1892]. [Untitled. Introduced by: "M. C. Emery, de Bologne, envoie les diagnoses de cinq nouveaux genres de Formicides".]. Bulletin Bimensuel de la Société Entomologique de France, 1892, cclxxv–cclxxvii.

Emery, C. (1893b) Voyage de M. E. Simon aux îles Philippines (mars et avril 1890). Formicides. Annales de la Société Entomologique de France, 62, 259–270.

Emery, C. (1895a) Beiträge zur Kenntniss der nordamerikanischen Ameisenfauna. (Schluss). Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere, 8, 257–360.

Emery, C. (1895b) Descriptions de quelques fourmis nouvelles d'Australie. Annales de la Société Entomologique de Belgique, 39, 345–358.

Emery, C. (1895c) Voyage de M. E. Simon dans l'Afrique australe (janvier-avril 1893). 3e mémoire. Formicides. Annales de la Société Entomologique de France, 64, 15–56.

Emery, C. (1895d) Die Gattung Dorylus Fab. und die systematische Eintheilung der Formiciden. Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere, 8, 685–778.

Emery, C. (1896) Clef analytique des genres de la famille des Formicides, pour la détermination des neutres. Annales de la Société Entomologique de Belgique, 40, 172–189.

Emery, C. (1897a) Viaggio di Lamberto Loria nella Papuasia orientale. XVIII. Formiche raccolte nella Nuova Guinea dal Dott. Lamberto Loria. Annali del Museo Civico di Storia Naturale di Genova, 38, 546–576.

Emery, C. (1897b) Formiche raccolte da Don Eugenio dei Principi Ruspoli, durante l’ultimo suo viaggio nelle regioni dei Somali e dei Galla. Annali del Museo Civico di Storia Naturale di Genova, 38, 596–605.

Emery, C. (1900a) Formicidarum species novae vel minus cognitae in collectione Musaei Nationalis Hungarici quas in Nova-Guinea, colonia germanica, collegit L. Biró. Publicatio secunda. Természetrajzi Füzetek, 23, 310–338.

Emery, C. (1900b) Formiche raccolte da Elio Modigliani in Sumatra, Engano e Mentawei. [part]. Annali del Museo Civico di Storia Naturale di Genova, 40 [= 2 (20)], 661–688.

Emery, C. (1901) Notes sur les sous-familles des Dorylines et Ponérines (Famille des Formicides). Annales de la Société Entomologique de Belgique, 45, 32–54.

Emery, C. (1902) Note mirmecologiche. Rendiconti delle Sessioni della Reale Accademia delle Scienze dell'Istituto di Bologna (n.s.), 6, 22–34.

Emery, C. (1909) Beiträge zur Monographie der Formiciden des paläarktischen Faunengebietes. (Hym.) Teil VIII. Deutsche Entomologische Zeitschrift, 1909, 355–376.

Emery, C. (1911) Hymenoptera. Fam. Formicidae. Subfam. Ponerinae. Genera Insectorum, 118, 1–125.

Emery, C. (1915) Formiche raccolte nell'Eritrea dal Prof. F. Silvestri. Bollettino del Laboratorio di Zoologia Generale e Agraria della Reale Scuola Superiore d'Agricoltura. Portici, 10, 3–26

Emery, C. & A. Forel. (1879) Catalogue des Formicides d'Europe. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 5, 441–481.

Escoubas, P., Whitman, D.W., Clement, J.L. & Blum, M.S. (1987) Larval appendages of two termitophagous ants, Hypoponera eduardi and Ponera pennsylvatica (Hymenoptera: Formicidae). Sociobiology, 13 (3), 241–247.

Esquivel, D.M.S., Wajnberg, E., Cernicchiaro, G.R. & Alves, O.C. (2004) Comparative magnetic measurements of migratory ant and its only termite prey. Journal of Magnetism and Magnetic Materials, 278, 117–121.

Evans, H.C. & Leston, D. (1971) A ponerine ant (Hym. Formicidae) associated with Homoptera on cocoa in Ghana. Bulletin of Entomological Research, 61, 357–362.

Fabricius, J.C. (1793) Entomologia systematica emendata et aucta. Secundum classes, ordines, genera, species, adjectis synonimis, locis observationibus, descriptionibus. Tome 2. C. G. Proft, Hafniae [= Copenhagen], 519 pp.

Fabricius, J.C. (1798) Supplementum entomologiae systematicae. Proft and Storch, Hafniae [= Copenhagen], 572 pp.

Fabricius, J.C. (1804) Systema Piezatorum secundum ordines, genera, species, adjectis synonymis, locis, observationibus, descriptionibus. C. Reichard, Brunswick, xiv + 15–439 + 30 pp.

Fales, H.M., Jones, T.H., Jaouni, T., Blum, M.S. & Schmidt, J.O. (1992) Phenylaklenals in ponerine (Leptogenys sp.) and myrmicine (Pogonomyrmex sp.) ants. Journal of Chemical Ecology, 18 (6), 847–854.

Fernandes, I.O., De Oliveira, M.L. & Delabie, J.H.C. (2014) Description of two new species in the Neotropical Pachycondyla foetida complex (Hymenoptera: Formicidae: Ponerinae) and taxonomic notes on the genus. Myrmecological News, 19, 133–163.

Ferrara, F., Maschwitz, U., Steghaus-Kovac, S. & Taiti, S. (1987) The genus Exalloniscus Stebbing, 1911 (Crustacea, Oniscidea) and its relationship with social insects. Pubblicazioni dell’Instituto entomol. Universita di Pavia, 36, 43–46.

Ferreira, R.S., Poteaux, C., Delabie, J.H.C., Fresneau, D. & Rybak, F. (2010) Stridulations reveal cryptic speciation in Neotropical sympatric Ants. PloS ONE, 5 (12), e15363.

Figueira, C.R.F. & Mathias, M.I.C. (2002) Histological, histochemical and morphometric study of female corpora allata of Pachycondyla striata ants (Hymenoptera: Ponerinae). Sociobiology, 39 (1), 77–87.

Fisher, B.L. (1997) Biogeography and ecology of the ant fauna of Madagascar (Hymenoptera: Formicidae). Journal of Natural History, 31, 269–302.

Fisher, B.L. (1999) Improving inventory efficiency: a case study of leaf-litter ant diversity in Madagascar. Ecological Applications, 9 (2), 714–731.[0714:iieacs];2

Fisher, B.L. (2006) Boloponera vicans gen.n. and sp.n. and two new species of the Plectroctena genus group (Hymenoptera: Formicidae). Myrmecologische Nachrichten, 8, 111–118.

Fisher, B.L. & Smith, M.A. (2008) A revision of Malagasy species of Anochetus Mayr and Odontomachus Latreille (Hymenoptera: Formicidae). PLoS ONE, 3 (5), e1787, 1–23.

Fletcher, D.J.C. (1973) ‘Army ant’ behaviour in the Ponerinae: a re-assessment. Proceedings of the VIIth international congress of the IUSSI, London, pp. 116–121.

Foutzik, S., Heinze, J., Oberstadt, B. & Herbers, J.M. (2002) Mate guarding and alternative reproductive tactics in the ant Hypoponera opacior. Animal Behaviour, 63, 597–604.

Forel, A. (1887) Fourmis récoltées à Madagascar par le Dr. Conrad Keller. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 7, 381–389.

Forel, A. (1890) Aenictus-Typhlatta découverte de M. Wroughton. Nouveaux genres de Formicides. Annales de la Société Entomologique de Belgique, 34, cii–cxiv.

Forel, A. (1891) Les Formicides. [part]. In: Grandidier, A. (Ed.), Histoire physique, naturelle, et politique de Madagascar. Volume XX. Histoire naturelle des Hyménoptères. Deuxième partie (28e fascicule). Hachette et Cie, Paris, v + 237 pp.

Forel, A. (1892) Nouvelles espèces de Formicides de Madagascar (récoltées par M. Sikora). Première série. Annales de la Société Entomologique de Belgique, 36, 516–535.

Forel, A. (1893a) Sur la classification de la famille des Formicides, avec remarques synonymiques. Annales de la Société Entomologique de Belgique, 37, 161–167.

Forel, A. (1893b) Formicides de l'Antille St. Vincent, récoltées par Mons. H. H. Smith. Transactions of the Entomological Society of London, 1893, 333–418.

Forel, A. (1895) A fauna das formigas do Brazil. Boletim do Museu Paraense de Historia Natural e Ethnographia, 1, 89–139.

Forel, A. (1897) Ameisen aus Nossi-Bé, Majunga, Juan de Nova (Madagaskar), den Aldabra-Inseln und Sansibar, gesammelt von Herrn Dr. A. Voeltzkow aus Berlin. Mit einem Anhang über die von Herrn Privatdocenten Dr. A. Brauer in Marburg auf den Seychellen und von Herrn. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 21, 185–208.

Forel, A. (1899) Formicidae. [part]. Biologia Centrali-Americana Hym, 3, 1–24.

Forel, A. (1900a) Ponerinae et Dorylinae d'Australie récoltés par MM. Turner, Froggatt, Nugent, Chase, Rothney, J.-J. Walker, etc. Annales de la Société Entomologique de Belgique, 44, 54–77.

Forel, A. (1900b) Les Formicides de l'Empire des Indes et de Ceylan. Part VI. Journal of the Bombay Natural History Society, 13, 52–65.

Forel, A. (1901a) I. Fourmis mexicaines récoltées par M. le professeur W.-M. Wheeler. II. A propos de la classification des fourmis. Annales de la Société Entomologique de Belgique, 45, 123–141.

Forel, A. (1901b) Variétés myrmécologiques. Annales de la Société Entomologique de Belgique, 45, 334–382.

Forel, A. (1901c) Nouvelles espèces de Ponerinae. (Avec un nouveau sous-genre et une espèce nouvelle d'Eciton). Revue Suisse de Zoologie, 9, 325–353.

Forel, A. (1905) Ameisen aus Java. Gesammelt von Prof. Karl Kraepelin 1904. Mitteilungen aus dem Naturhistorischen Museum in Hamburg, 22, 1–26.

Forel, A. (1908) Fourmis de Costa-Rica récoltées par M. Paul Biolley. Bulletin de la Société Vaudoise des Sciences Naturelles, 44, 35–72.

Forel, A. (1912) Descriptions provisoires de genres, sous-genres, et espèces de Formicides des Indes orientales. Revue Suisse de Zoologie, 20, 761–774.

Forel, A. (1913a) Fourmis de Rhodesia, etc. récoltées par M. G. Arnold, le Dr. H. Brauns et K. Fikendey. Annales de la Société Entomologique de Belgique, 57, 108–147.

Forel, A. (1913b) Formicides du Congo Belge récoltés par MM. Bequaert, Luja, etc. Revue Zoologique Africaine (Brussels), 2, 306–351.

Forel, A. (1915a) Fauna Simalurensis. Hymenoptera Aculeata, Fam. Formicidae. Tijdschrift voor Entomologie, 58, 22–43.

Forel, A. (1915b) Results of Dr. E. Mjöbergs Swedish Scientific Expeditions to Australia 1910-13. 2. Ameisen. Arkiv för Zoologi, 9 (1) 16, 1–119.

Forel, A. (1917) Cadre synoptique actuel de la faune universelle des fourmis. Bulletin de la Société Vaudoise des Sciences Naturelles, 51, 229–253.

Forel, A. (1928) The social world of the ants compared with that of man. Vol. II. (tr. C. K. Ogden). G. P. Putnam’s Sons, Ltd., London, xx + 444 pp.

Fourcassié, V., Henriques, A. & Fontella, C. (1999) Route fidelity and spatial orientation in the ant Dinoponera gigantea (Hymenoptera, Formicidae) in a primary forest: a preliminary study. Sociobiology, 34 (3), 505–524.

Fourcassié, V. & Oliveira, P.S. (2002) Foraging ecology of the giant Amazonian ant Dinoponera gigantea (Hymnoptera, Formicidae, Ponerinae): activity schedule, diet and spatial foraging patterns. Journal of Natural History, 36, 2211–2227.

Fowler, H.G. (1980) Populations, prey capture and sharing, and foraging of the Paraguayan ponerine Odontomachus chelifer Latreille. Journal of Natural History, 14 (1), 79–84.

Fowler, H.G. (1985) Populations, foraging and territoriality in Dinoponera australis (Hymenoptera, Formicidae). Revista Brasileira de Entomologia, 29, 443–447.

Freitas, A.V.L. (1995) Nest relocation and prey specialization in the ant Leptogenys propefalcigera Roger (Formicidae: Ponerinae) in an urban area in southeastern Brazil. Insectes Sociaux, 42, 453–456.

Fresneau, D. (1984) Développement ovarien et status social chez une fourmi primitive Neoponera obscuricornis Emery (Hym. Formicidae, Ponerinae). Insectes Sociaux, 31 (4), 387–402.

Fresneau, D. (1985) Individual foraging and path fidelity in a ponerine ant. Insectes Sociaux, 32 (2), 109–116.

Fresneau, D. & Dupuy, P. (1988) A study of polyethism in a ponerine ant: Neoponera apicalis (Hymenoptera, Formicidae). Animal Behaviour, 36, 1389–1399.

Fukumoto, Y. & Abe, T. (1983) Social organization of colony movement in the tropical ponerine ant, Diacamma rugosum (Le Guillou). Journal of Ethology, 1, 101–108.

Fukumoto, Y., Abe, T. & Taki, A. (1989) A novel form of colony organization in the “queenless” ant Diacamma rugosum. Physiology and Ecology Japan, 26, 55–61.

Fukuzawa, M., Arakura, F., Yamazaki, Y., Uhara, H. & Saida, T. (2002) Urticaria and anaphylaxis due to sting by an ant (Brachyponera chinensis). Acta Dermato Venereologica, 82 (1), 59.

Ganeshaiah, K.N. & Veena, T. (1991) Topology of the foraging trails of Leptogenys processionalis – why are they branched? Behavioral Ecology and Sociobiology, 29, 263–270.

Garcia-Pérez, J.A., Blanco-Piñon, A., Mercado-Hernández, R. & Badii, M. (1997) El comportamiento depredador de Pachycondyla harpax Fabr. sobre Gnathamitermes tubiformans Buckley en condiciones de cautiverio. Southwestern Entomology, 22 (3), 345–353.

Giovannotti, M. (1996) The stridulatory organ of five Ponerinae species: a SEM study (Hymenoptera, Formicidae). Fragmenta Entomologica, 28, 157–165.

Gobin, B., Heinze, J., Strätz, M. & Roces, F. (2003a) The energetic cost of reproductive conflicts in the ant Pachycondyla obscuricornis. Journal of Insect Physiology, 49, 747–752.

Gobin, B., Ito, F. & Billen, J. (2003b) The subepithelial gland in ants: a novel exocrine gland closely associated with the cuticle surface. Acta Zoologica, 84, 285–291.

Gobin, B., Ito, F., Billen, J. & Peeters, C. (2008) Degeneration of sperm reservoir and the loss of mating ability in worker ants. Naturwissenschaften, 95 (11), 1041–1048.

Gobin, B., Ito, F., Peeters, C. & Billen, J. (2006) Queen-worker differences in spermatheca reservoir of phylogenetically basal ants. Cell Tissue Research, 326, 169–178.

Gopinath, A., Gadagkar, R. & Rao, M.R.S. (2001) Identification of polymorphic microsatellite loci in the queenless, ponerine ant Diacamma ceylonense. Molecular Ecology Notes, 1, 126–127.

Goss, S., Fresneau, D., Deneubourg, J.L., Lachaud, J.-P. & Valenzuela-Gonzalez, J. (1989) Individual foraging in the ant Pachycondyla apicalis. Oecologia, 80, 65–69.

Gotoh, A. & Ito, F. (2008) Seasonal cycle of colony structure in the ponerine ant Pachycondyla chinensis in western Japan (Hymenoptera, Formicidae). Insectes Sociaux, 55, 98–104.

Gotoh, A., Sameshima, S., Tsuji, K., Matsumoto, T. & Miura, T. (2005) Apoptotic wing degeneration and formation of an altriusm-regulating glandular appendage (gemma) in the ponerine ant Diacamma sp. from Japan (Hymenoptera, Formicidae, Ponerinae). Development Genes and Evolution, 2215, 69–77.

Gotwald, W.H. Jr. & Brown, W.L. Jr. (1967) [1966] The ant genus Simopelta (Hymenoptera: Formicidae). Psyche (Camb.), 73, 261–277.

Gronenberg, W. (1995a) The fast mandible strike in the trap-jaw ant Odontomachus. I. Temporal properties and mor


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