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A new species of anchovy, Encrasicholina macrocephala (Clupeiformes: Engraulidae), from the northwestern Indian Ocean



A new species of anchovy, Encrasicholina macrocephala, is described on the basis of 17 specimens collected from the Red Sea and the Arabian Sea. The species is closely related to E. devisi in that both species have three unbranched rays in the dorsal and anal fins, and a long upper jaw (posterior tip extending beyond posterior margin of preopercle). However, the new species is distinguished from E. devisi in having lower counts of total pectoral-fin rays (11–14 vs. 13–15) and pseudobranchial filaments (15–18 vs. 18–22), longer head length (29.5–31.7% of standard length vs. 25.4–28.9% in the latter), upper-jaw length (21.3–23.5% vs. 14.6–21.3%), and lower-jaw length (19.5–21.2% vs. 14.2–19.5%), and a shorter distance between the dorsal-fin origin to the pectoral-fin insertion (83.2–95.2% of head length vs. 97.6–126.1%).



Teleostei, new species, Encrasicholina devisi, Red Sea, Arabian Sea

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Hardenberg, J.D.F. (1933a) Notes on some genera of the Engraulidae. Natuurkundig Tijdschrift voor Nederlandsch Indië, 93 (2), 230–257.

Hardenberg, J.D.F. (1933b) New Stolephorus species of the Indo-Australian seas. Natuurkundig Tijdschrift voor Nederlandsch Indië, 93 (2), 258–263.

Hardenberg, J.D.F. (1934) Some remarks on the genus Stolephorus Lacépède in the Indo-Australian Archipelago. Treubia Buitenzorgë, 14 (3), 313–375.

Hata, H., Motomura, H. & Ishimori, H. (2012) First Japanese record of an engraulid fish, Encrasicholina devisi (Clupeiformes), collected from Kagoshima Prefecture, southern Japan and comparisons with congeners. Japanese Journal of Ichthyology, 59 (2), 125–134.

Kimura, S., Hori, K. & Shibukawa, K. (2009) A new anchovy, Stolephorus teguhi (Clupeiformes: Engraulidae), from North Sulawesi, Indonesia. Ichthyological Research, 56, 292–295.

Sabaj Pérez, M.H. (Ed.) (2010) Standard symbolic codes for institutional resource collections in herpetology and ichthyology: an online reference. Version 2.0 (8 November 2010). Electronically accessible. American Society of Ichthyologists and Herpetologists, Washington, DC. Available from: (accessed 3 March 2015)

Strasburg, D.W. (1960) A new Hawaiian engraulid fish. Pacific Science, 14 (No. 4), 395–399.

Whitehead, P.J.P., Nelson, G.J. & Wongratana, T. (1988) FAO species catalogue Vol. 7. Clupeoid Fishes of the World (Suborder Clupeoidei). An Annotated and Illustrated Catalogue of the Herrings, Sardines, Pilchards, Sprats, Shads, Anchovies and Wolf-herrings. Part 2 – Engraulidae. FAO Fisheries Synopsis, No. 125, 7 (pt. 2), i–viii + 305–579.

Wongratana, T. (1983) Diagnoses of 24 new species and proposal of a new name for a species of Indo-Pacific clupeoid fishes. Japanese Journal of Ichthylogy, 29 (4), 385–407.

Wongratana, T., Munroe, T.A. & Nizinski, M.S. (1999) Order Clupeiformes. Engraulidae. Anchovies. In: Carpenter, K.E. & Niem, V.H. (Eds.), FAO Species Identification Guide for Fishery Purposes. The Living Marine Resources of the Western Central Pacific. Vol. 3. Batoid Fishes, Chimaeras and Bony Fishes Part 1 (Elopidae to Linophrynidae). FAO, Rome, pp. 1698–1753.


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