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Garra robertsi, a new cyprinid (Cypriniformes: Cyprinidae) fish species from Borneo



Garra robertsi is described from specimens collected from the Sungai Bongan and Tempassuk rivers in Sabah, Borneo. The species is differentiated from G. borneensis, its only congener on the island of Borneo, in having five (versus four) transverse scale rows above lateral line, the first branched dorsal-fin ray extending beyond the posterior-most extent of any other part of the dorsal fin when depressed (versus not extending posteriorly beyond last ray when depressed), breast with deeply embedded scales (versus exposed scales), fewer tubercles on snout, thin (versus thick) anteromedial fold on the lower lip, absence (versus presence) of a lateral stripe, absence (versus presence) of a stark, contrasting black stripe on lower caudal-fin rays, and other pigmentation characteristics.



Garrinae, Malaysia, Sungai Bongan River, Tempassuk River

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Eschmeyer, W.N. (Ed.) (1998) Catalog of Fishes. Vol. 1–3. Center for Biodiversity Researchand Information. Special Publication 1. California Academy of Sciences, KNI Incorporated, Anaheim, California, 2905 pp.

Eschmeyer, W.N. (Ed.) (2015) Catalog of fishes: genera, species, references. Electronic version. Available from: (accessed 28 January 2015)

Kottelat, M. (2013) The fishes of the inland waters of southeast Asia: a catalogue and core bibliography of the fishes known to occur in freshwaters, mangroves and estuaries. The Raffles Bulletin of Zoology, 27 (Supplement), 1–663.

Kullander, S.O. & Fang, F. (2004) Seven new species of Garra (Cyprinidae:Cyprininae) from the Rakhine Yoma, southern Myanmar. Ichthyological exploration of freshwaters, 15 (3), 257–278.

Nebeshwar, K., Vishwanath, W. & Das, D.N. (2009) Garra arupi, a new cyprinid fish species (Cypriniformes: Cyprinidae) from upper Brahmaputra basin in Arunachal Pradesh, India. Journal of Threatened Taxa, 1 (4), 197–202.

Sabaj Pérez, M.H. (Ed.) (2014) Standard symbolic codes for institutional resource collections in herpetology and ichthyology: an Online Reference. Version 5.0 (22 September 2014). American Society of Ichthyologists and Herpetologists, Washington, DC. Electronically accessible. Available from: (accessed 23 June 2015)

Yang, L., Arunachalam, M., Sado, T., Levin, B.A., Golubtsov, A.S., Freyhof, J., Friel, J.P., Chen, W.-J., Hirt, M.V., Manickam, R., Agnew, M.K., Simons, A.M., Saitoh, K., Miya, M., Mayden, R.L. & He, S. (2012) Molecular phylogeny of the cyprinid tribe Labeonini (Teleostei: Cypriniformes). Molecular Phylogenetics and Evolution, 65 (2012), 362–379.

Vaillant, L.L. (1902) Résultats zoologiques de l'expédition scientifique Néerlandaise au Bornéo central. Poissons. Notes from the Leyden Museum, 24 (Note 1), 1–166, pls. 1–2.


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