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A review of the genera Cleigastra Macquart, Gonarcticus Becker, Gonatherus Rondani, Hexamitocera Becker, Nanna Strobl, Orthacheta Becker and Spathephilus Becker (Diptera, Scathophagidae) of Russia



Flies of the genera Cleigastra Macquart, 1835, Gonarcticus Becker, 1894, Gonatherus Rondani, 1856, Hexamitocera Becker, 1894, Nanna Strobl, 1894, Orthacheta Becker, 1894 and Spathephilus Becker, 1894 (all Scathophagidae) of the fauna of Russia are reviewed. Key to genera, generic descriptions and keys for determination of species are given, and data on geographical distribution are summarized. One species, Nanna cryophila sp. nov., is described as new to science. One new synonymy is proposed: Nanna kamtschatkense (Hendel, 1930) = Nanna tibiella (Zetterstedt, 1838). Orthacheta cornuta (Loew, 1863) is recorded from Europe for the first time. Gonarcticus arcticus (Becker, 1907) is newly recorded from the Palaearctic Region and Russia. Additionally, Nanna flavipes (Fallén, 1819) is newly recorded from China, and Spathephilus nigriventris (Loew, 1864) is newly recorded from Kazakhstan.



Diptera, dung flies, new records, new species, Russia, review, Scathophagidae

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Aldrich, J.M. (1905) A catalogue of North American Diptera (or two-winged flies). Smithsonian miscellaneous collections, 46 (1444), 1–680.

Barták, M. & Kubík, Š. (2013) A redefinition of Micropselapha Becker, 1894 (Diptera, Scathophagidae) with description of a new species from Central Europe. Entomological News, 123 (2), 101–109.

Becker, T. (1894) Dipterologische Studien. I. Scatomyzidae. Berliner Entomologische Zeitschrift, 39 (1), 77–196.

Becker, T. (1907) Beschreibung von 3 neuen Dipteren aus Ost-Grönland. Meddelelser om Grönland, 29, 411–414.

Becker, T. (1915) Cyclorrhapha schizophora. Holometopa. In: Becker, Th., Dziedzicki, H., Schnabl, J. & Villeneuve, J.P. (Eds.), Résultats scientifiques de l'Expédition des frères Kuznecov (Kouznetzov) à l'Oural Arctique en 1909, sous la direction de H. Baklund. Livr.7. Diptera. Mémoires de l'Académie Impériale des Sciences, Classe Physico-Mathématique, Série 8, 28 (7), pp. 63–67.

Cumming, J.M. & Wood, D.M. (2009) Adult morphology and terminology. In: Brown, B.V., Borkent, A., Cumming, J.M., Wood, D.M., Woodley, N.E. & Zumbado, M.A. (Eds.), Manual of Central American Diptera. Vol. 1. National Research Council Press, Ottawa, pp. 9–50.

Engelmark, R. (1999) Dungflies (Diptera: Scathophagidae) collected by the Swedish-Russian tundra ecology expedition '94, with the description of two new species; Nanna indotatum and Cochliarium sibiricum. Entomologisk Tidskrift, 120 (4), 157–167.

Fallén, C.F. (1819) Scatomyzides. Lundae, Berling, 10 pp.

Fallén, C.F. (1826) Supplementum Dipterorum Sveciae. Berling, Lundae, 16 pp.

Ferrar, P. (1987) A guide to the breeding habits and immature stages of Diptera Cyclorrhapha. Entomonograph, 8 (1–2), 1–907.

Gorodkov, K.B. (1970) Family Scathophagidae (Cordyluridae, Scatomyzidae, Scopeumatidae). In: Bei-Bienko, G.Y. (Ed.), Key to the insects of the European part of the USSR, V. Diptera, Siphonaptera, 2, pp. 440–458. [in Russian]

Gorodkov, K.B. (1986) Family Scathophagidae. In Soós, Á. & Papp, L. (Eds), Catalogue of Palaearctic Diptera. Vol.11. Scathophagidae-Hypodermatidae. Akadémiai Kiadó, Budapest, pp. 11–41.

Grochowska, M. (2006) Remarks on the morphology and biology of Cleigastra apicalis (Meigen, 1826) (Diptera, Scathophagidae). Acta Zoologica (Stockholm), 87, 247–252.

Groth, U. (1969) Zur Entwicklung und Biologie von Cnemopogon apicalis Wied. (Diptera: Cordyluridae). Wissenschaftliche Zeitschrift der Ernst-Moritz-Arndt-Universität Greifswald, Mathematisch-Naturwissenschaftliche Reihe, 18 (1/2), 85–92.

Hackman, W. (1956) The Scatophagidae (Dipt.) of Eastern Fennoscandia. Societas pro Fauna et Flora Fennica, Fauna fennica II. 67 pp. [Helsingforsiae]

Hadley, A. (2007) CombineZM. Open source image processing software package for creating extended depth of field images. Available from: (accessed 17 January 2011)

Humala, A.E. & Polevoi, A.V. (2009) On the Insect fauna of south-east Karelia. Trudi Karelskogo nauchnogo centra RAN, 4, 53–75.

Jong, H. de (2000) Family Scathophagidae. In: Papp, L. & Darvas, B. (Eds.), Contributions to a Manual of Palaearctic Diptera. Appendix. Science Herald, Budapest, pp. 431–445.

Kutty, S.N., Bernasconi, M.V., Šifner F. & Meier, R. (2007) Sensitivity analysis, molecular systematics and natural history evolution of Scathophagidae (Diptera: Cyclorrhapha: Calyptratae). Cladistics, 23, 64–83.

Loew, H. (1863) Diptera Americana septentrionalis indigena. Berliner Entomologische Zeitschrift, 7, 1–32.

Loew, H. (1864) Acht neue Cordylura-Arten. Wiener Entomologische Monatschrift, 8, 17–26.

Loew, H. (1869) Diptera Americana septentrionalis indigena. Berliner Entomologische Zeitschrift, 13, 129–186.

Loew, H. (1873) Beschreibungen europäischen Dipteren. Dritter Band. Systematische Beschreibung der bekannten europäischen zweiflügeligen Insecten von Johann Wilhelm Meigen. Zehnter Theil oder vierter Supplementband. Schmidt, Halle, 320 pp.

Macquart, J. (1835) Histoire naturelle des Insectes. Diptères. Vol. 2. Roret, Paris, 710 pp.

Malloch, J.R. (1920) XII. Descriptions of Diptera of the families Anthomyidae and Scatophagidae. The Ohio Journal of Science, 20 (7), 267–391.

McAlpine, J.F. (1981) Morphology and terminology – Adults. In: McAlpine, J.F., Peterson, B.V., Shewell, G.E., Teskey, H.J., Vockeroth, J.R. & Wood, D.M. (Eds.), Manual of Nearctic Diptera. Vol. 2. Agriculture Canada Research Branch, Monograph 27, Ottawa, pp. 9–64.

Meigen, J.W. (1826) Systematische Beschreibung der bekannten europäischen zweiflügeligen Insekten. Siebenter Theil oder Supplementband. Schultz, Hamm, 412 pp.

Meigen, J.W. (1838) Systematische Bescreibung der bekannten europäischen zweiflügeligen Dipteren. Siebenter Theil oder Supplementband. Schultz, Hamm, 434 pp.

Merriam Webster (1997) Merriam Webster’s Geographical Dictionary. Third Edition. Springfield, MA, 1361 pp.

Neff, S.E. & Wallace, J.B. (1969) Biology and Description of Immature Stages of Orthacheta hirtipes, a Predator of Cordilura spp. Annals of the Entomological Society of America, 62 (4), 785–790.

Ovchinnikov, A.N. (2004) Data on the fauna of Scathophagidae (Diptera) of the Volga Region. Russian Entomological Journal, 12 (4) 421–423. [2003]

Ovchinnikov, A.N. (2009) The Ovipositor Morphology in the Members of the Family Scathophagidae (Diptera) with Reference to Their Biology. Entomological Review, 89 (6), 623–636.

Ozerov, A.L. (2007) On Scathophagidae species (Insecta: Diptera), described by F. Hendel from the materials of the Swedish Kamchatka expedition 1920–1922. Far Eastern Entomologist, 174, 5–8.

Ozerov, A.L. (2008) Two new species of the genus Norellia Robineau-Desvoidy, 1830 (Diptera, Scathophagidae) from Altai. Far Eastern Entomologist, 184, 1–4.

Ozerov, A.L. (2009) New species of Scathophagidae (Diptera). Russian Entomological Journal, 17 (4), 419–427. [2008]

Ozerov, A.L. (2010a) On the Palaearctic fauna of Norellisoma Wahlgren, 1917 (Diptera: Scathophagidae). Russian Entomological Journal, 18 (3), 229–234. [2009]

Ozerov, A.L. (2010b) A review of Palaearctic species of the genera Delina Robineau-Desvoidy, 1830 and Neochirosia Malloch, 1917 (Insecta: Diptera: Scathophagidae). Russian Entomological Journal, 18 (3), 235–241. [2009]

Ozerov, A.L. (2010c) Flies of the genus Parallelomma Becker in Strobl, 1894 (Diptera, Scathophagidae) in the fauna of Russia. Russian Entomological Journal, 18 (4), 309–317. [2009, in Russian and English]

Ozerov, A.L. (2010d) A Review of Palaearctic species of the genus Nanna Strobl, 1894 (Diptera, Scathophagidae) with shining stripes or spots on scutum. Far Eastern Entomologist, 214, 1–8.

Ozerov, A.L. (2010e) Five new species of Scathophagidae (Diptera) from Russia. Russian Entomological Journal, 19 (2), 157–166.

Ozerov, A.L. (2012) A review of World species of the genus Spaziphora Rondani, 1856 (Diptera, Scathophagidae). Russian Entomological Journal, 20 (4), 423–430. [2011]

Ozerov, A.L. (2013) Description of five new species and notes on taxonomy of Scathophagidae (Diptera). Russian Entomological Journal, 22 (1), 81–89.

Ozerov, A.L. (2014) Notes on types of three species of Scathophagidae (Diptera) described by I.W. Zetterstedt. Russian Entomological Journal, 23 (4), 317–320.

Ozerov, A.L. & Krivosheina, M.G. (2011) The first record of Norellia tipularia (Fabricius, 1794) (Diptera, Scathophagidae) for the fauna of Russia. Russian Entomological Journal, 20 (1), 101–103.

Ozerov, A.L. & Krivosheina, M.G. (2012a) Revision of dung flies of the genus Mixocordylura Hendel (Diptera, Scathophagidae), with the description of new species from Altay. Russian Entomological Journal, 20 (4), 431–435. [2011]

Ozerov, A.L. & Krivosheina, M.G. (2012b) To knowledge of the genus Cordilura Fallén (Diptera, Scathophagidae), with description of a new species from the Russian Far East. Far Eastern Entomologist, 249, 1–4.

Ozerov, A.L. & Krivosheina, M.G. (2012c) A review of World species of the genus Pleurochaetella Vockeroth, 1965 (Diptera, Scathophagidae). Russian Entomological Journal, 21 (4), 435–440.

Ozerov, A.L. & Krivosheina, M.G. (2013) New and little known palaearctic species of Scathophagidae (Diptera). Far Eastern Entomologist, 270, 1–6.

Ozerov, A.L. & Krivosheina, M.G. (2014a) A review of species of the genera Bostrichopyga Becker, 1894 and Paracosmetopus Hackman, 1956 (Diptera, Scathophagidae). Russian Entomological Journal, 23 (1), 85–90.

Ozerov, A.L. & Krivosheina, M.G. (2014b) To the fauna of dung flies (Diptera: Scathophagidae ) of Russian Far East. Russian Entomological Journal, 23 (3), 203–222.

Pape, T., Blagoderov, V. & Mostovski, M.B. (2011) Order Diptera Linnaeus, 1758. In: Zhang, Z.-Q., (Ed.), Animal biodiversity: an outline of higher-level classification and survey of taxonomic richness. Zootaxa, 3148, 222–229.

Ringdahl, O. (1936) Anteckningar till svenska arter av familjen Scopeumatidae (Diptera). Entomologisk Tidskrift, 57 (2–3), 158–179. [in Swedish]

Rondani, C. (1856) Dipterologiae Italicae Prodromus. Vol. I. Genera Italica ordinis dipterorum ordinatim disposita et distincta et in familias et stirpes aggregata. A. Stoschi, Parmae, 228 pp.

Roser, K.L.F. von (1840) III. Beiträge zur Vaterlandskunde Erster Nachtrage zu dem im Jahre 1834 bekannte gemachten Verziechnisse in Württemberg vorkommender zweiflügliger Insekten. Correspondenzblatt der Landwirtschaft aus dem Vereins in Württemburg, 1 (Supplement 1), 49–64. [Stuttgart]

Šifner, F. (2003) The family Scathophagidae (Diptera) of the Czech and Slovak Republics (with notes on selected Palaearctic taxa). Acta Musei Nationalis Pragae, Series B, Natural History, 59 (1–2), 1–90.

Šifner, F. (2008) A catalogue of the Scathophagidae (Diptera) of the Palaearctic region, with notes on their taxonomy and faunistics. Acta Entomologica Musei Nationalis Pragae, 48 (1), 111–196.

Sorokina, V.S. & Pont, A.C. (2014) The localities of Arctic Diptera (Insecta) collected by the Russian Kara Expedition of 1909. Journal of Natural History, 49 (25–26), pp. 1585–1598.

Strobl, G. (1894) Die Dipteren von Steiermark. II. Theil. Mitteilungen des Naturwissenschaftlichen Vereines für Steiermark, 30, 1–152.

Stuckenberg, B.R. (1999) Antennal evolution in the Brachycera (Diptera), with a reassessment of terminology relating to the flagellum. Studia Dipterologica, 6, 33–48.

Vockeroth, J.R. (1965) Subfamily Scatophaginae. In: Stone, A., Sabrosky, C.W., Wirth, W.W., Foote, R.H. & Culson, J.R. (Eds.), A catalog of the Diptera of America north of Mexico. United States Department of Agriculture, Agriculture Handbook 276, pp. 826–842.

Vockeroth, J.R. (1987) 103. Scathophagidae. In: McAlpine, J.F., Peterson, B.V., Shewell, G.E., Teskey, H.J., Vockeroth, J.R. & Wood, D.M. (Eds.), Manual of Nearctic Diptera. Vol. 2. Agriculture Canada Research Branch, Monograph 28, Ottawa, pp. 1085–1097.

Vockeroth, J.R. (1995) Validation of nomina nuda of Nearctic Tethinidae, Scathophagidae, and Muscidae proposed in Manual of Nearctic Diptera. Proceedings of the Entomological Society of Washington, 93 (3), 732–734.

Westwood, J.O. (1840) An introduction to the modern classication of insects. Synopsis of the genera of British Insects. Longman, Orme, Brown, Green and Longmans, London, 158 pp.

Zetterstedt, J.W. (1838) Sectio tertia. Diptera. In: Insecta Lapponica. Leopold Voss, Lipsiae [= Leipzig], pp. 477–868. [‘1840’]

Zetterstedt, J.W. (1846) Diptera Scandinaviae disposita et descripta. Tomus quintus. Ex officina lundbergiana, Lundae [= Lund], 1739–2162.


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