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There are recent faunistic surveys of selected insect taxa (e.g. Mutillidae, Scarabaeoidea, and Tenebrionidae) in Wisconsin but a formal investigation of the Meloidae (blister beetles) is lacking. The blister beetle fauna of several states has been published, but this study represents the first in the Midwestern United States. We provide a comprehensive list of all meloid species documented from Wisconsin. Also included are taxonomic keys as well as summaries for each species (species pages) which includes taxonomy, description, and natural history. Specimens were obtained from public and private collections, and field sampling. This survey advances our knowledge of meloids in Wisconsin as well as provides a contribution beyond this geographic area. During this survey, 28 meloid species in seven genera were documented in Wisconsin from 2605 specimens. Of these taxa, 10 species are considered new state records. While Epicauta pensylvanica represented nearly half of the specimens reviewed, and likely inhabits all counties within the state, other species were rarely encountered. This includes 10 species which were represented by seven specimens or fewer in this study. It is unclear if the rarity of these specimens is correlated with the rarity of the species or if it is due to other factors. Regardless, these rarely collected meloids in Wisconsin warrant further attention.



Coleoptera, blister beetle, Meloidae, Wisconsin, distribution, natural history, taxonomy, checklist


Abdullah, M. (1964) The natural classification and new species of Pedilus Fischer (Coleoptera, Anthicidae, Pedilinae). Deutsche Entomologische Zeitschrift, 11, 145–174.

Adams, C.L. & Selander, R.B. (1979) The biology of blister beetles of the Vittata Group of the genus Epicauta (Coleoptera, Meloidae). Bulletin of the American Museum of Natural History, 162, 137–266.

Arnold, D.C. (1976) Blister beetles (Coleoptera: Meloidae) of Oklahoma. Oklahoma State University Agricultural Experiment Station Technical Bulletin, T-145, 1–68.

Baerg, W.J. (1925) Control measures for blister beetles. Arkansas Agricultural Experiment Station Bulletin, 201, 1–8.

Bahme, A.J. (1968) Cantharides toxicosis in the equine. Southwestern Vet, 21, 147–149.

Batra, S.W.T. (1980) Ecology, behavior, pheromones, parasites and management of the sympatric vernal bees Colletes inaequalis, C. thoracicus and C. validus. Journal of the Kansas Entomological Society, 53, 509–538.

Beauregard, H. (1890) Les insects vésicants. Paris, 544 pp.

Bilsing, S.W. (1920) Quantitative studies in the food of spiders. Ohio Journal of Science, 20, 215–260.

Blatchley, W.S. (1910) An illustrated descriptive catalogue of the Coleoptera of beetles (exclusive of Rhynchophora) known to occur in Indiana. Nature Publishing Company, Indianapolis, 1386 pp.

Blodgett, S.L., Carrel, J.E. & Higgins, R.A. (1991) Cantharidin content of blister beetles (Coleoptera: Meloidae) collected from Kansas alfalfa and implications for inducing cantharidiasis. Environmental Entomology, 20, 776–780.

Blodgett, S.L., Denke, P.M. & Knerr, V. (2010) Blister beetles of Montana. Montana State University Extension Service MT200209 AG, 100-210SA, 1–4.

Blodgett, S.L. & Higgins, R.A. (1988) Blister beetles (Coleoptera: Meloidae) in Kansas: historical perspective and results of an intensive alfalfa survey. Journal of Economic Entomology, 8, 1456–1462.

Bologna, M.A. (1991) Coleoptera Meloidae. Fauna d’Italia. XXVIII. Calderini, Bologna, 541 pp.

Bologna, M.A., Oliverio, M., Pitzalis, M. & Mariottini, P. (2008) Phylogeny and evolutionary history of the blister beetles (Coleoptera, Meloidae). Molecular Phylogenetics and Evolution, 48, 679–693.

Bologna, M.A. & Pinto, J.D. (1998) A review of the Afrotropical species of Meloe Linnaeus 1758 (Coleoptera, Meloidae) with descriptions of first instar larvae, a key to species and an annotated catalogue. Tropical Zoology, 11, 19–59.

Bologna, M.A. & Pinto, J.D. (2001) Phylogenetic studies of Meloidae (Coleoptera), with emphasis on the evolution of phoresy. Systematic Entomology, 26, 33–72.

Bologna, M.A., Turco, F. & Pinto, J.D. (2013) The Meloidae (Coleoptera) of Australasia: a generic review, descriptions of new taxa, and a challenge to the current definition of subfamilies posed by exceptional variation in male genitalia. Invertebrate Systematics, 27, 391–427.

Bomar, C.R. (1993) The blister beetles (Coleoptera: Meloidae) of Wyoming. University of Wyoming Agricultural Experiment Station, B-981, 1–34.

Bouseman, J.K. (1966) Gnathium minimum (Say) in Illinois (Coleoptera: Meloidae). The Coleopterists’ Bulletin, 20, 59.

Brimley, C. (1938) The Insects of North Carolina. North Carolina Department of Agriculture, Raleigh, North Carolina, 560 pp.

Burgess, L. (1983) Damage to rapeseed plants by two species of blister beetles (Coleoptera: Meloidae). Canadian Entomologist, 115, 875–876.

Butler, L. (1984) Additional observations on the association of Pedilus (Pedilidae) with Meloe (Coleoptera: Meloidae). Entomological News, 95, 101–102.

Capinera, J.L., Gardner, D.R. & Stermitz, F.R. (1985) Cantharidin levels in blister beetles (Coleoptera: Meloidae) associated with alfalfa in Colorado. Journal of Economic Entomology, 78, 1052–1055.

Carrel, J.E. (1999) Contrasting responses of southern house spiders and raccoons to blister beetle prey. Journal of Chemical Ecology, 25, 1295–1303.

Carrel, J.E. & Eisner, T. (1974) Cantharidin: Potent feeding deterrent to insects. Science, 183, 755–757.

Carruth, L.A. (1931) The Meloidae of South Dakota. Entomological News, 42, 50–55.

Chittenden, F.H. (1898) Insects injurious to beans and peas. USDA 1898 Yearbook, 233–260.

Chittenden, F.H. (1903) A brief account of the principal insect enemies of the sugar beet. USDA Division of Entomology Bulletin, 43, 1–71.

Christoffel, R., Hay, R. & Monroe, M. (2002) Turtles and lizards of Wisconsin. Publication ER.104. Wisconsin Department of Natural Resources, Madison, Wisconsin, 48 pp.

Christoffel, R., Hay, R. & Ramirez, L. (2000) Snakes of Wisconsin. Publication ER-100. Wisconsin Department of Natural Resources, Madison, Wisconsin, 32 pp.

Christoffel, R., Hay, R. & Wolfgram, M. (2001) Amphibians of Wisconsin. Publication ER-105. Wisconsin Department of Natural Resources, Madison, Wisconsin, 44 pp.

Church, N.S. & Gerber, G.H. (1977) Observations on the ontogeny and habits of Lytta nuttalli, L. viridana, and L. cyanipennis (Coleoptera: Meloidae): the adults and eggs. Canadian Entomologist, 109, 565–573.

Claypole, E.W. (1881) Notes on Lytta (blister beetles). Proceedings of the Entomological Society of Ontario, 31–32.

Cline, A.R. & Huether, J.P. (2011) Revision of the nearctic blister beetle genus Tricrania LeConte, 1860 (Coleoptera: Meloidae: Nemognathinae). Zootaxa, 2832, 1–43.

Coskey, R.J. (1984) Treatment of plantar warts in children with a salicylic acid-podophyllin-cantharidin product. Pediatric Dermatology, 2, 71–73.

Curtis, J.T. (1959) The Vegetation of Wisconsin. University of Wisconsin Press, Madison, 657 pp.

Dennis, D.S. & Lavigne, R.J. (1975) Comparative behavior of Wyoming robber flies II (Diptera: Asilidae). Agricultural Experiment Station University of Wyoming-Laramie Science Monograph, 30, 1–68.

Dillon, L.S. (1952) The Meloidae (Coleoptera) of Texas. American Midland Naturalist, 48, 330–420.

Dunford, J.C. & Young, D.K. (2004) An annotated checklist of Wisconsin darkling beetles (Coleoptera: Tenebrionidae) with comparisons to the western Great Lakes fauna. Transactions of the American Entomological Society, 130, 57–76.

Dunn, G.A. (1996) Insects of the Great Lakes Region. The University of Michigan Press, Ann Arbor, Michigan, 324 pp.

Eickwort, G.C. (1981) Agapostemon virescens (Hymenoptera: Halictidae; Halictinae) as a larval host of Lytta (Pomphopoea) sayi (Coleoptera: Meloidae). Journal of the Kansas Entomological Society, 54, 519–520.

Enns, W.R. (1956) A revision of the genera Nemognatha, Zonitis, and Pseudozonitis (Coleoptera, Meloidae) in American north of Mexico, with a proposed new genus. The University of Kansas Science Bulletin, 37, 685–909.

Erickson, E.H. (1973) First-stage larvae of Tegrodera erosa aloga and Gnathium obscurum (Coleoptera: Meloidae). Annals of the Entomological Society of America, 66, 785–787.

Erickson, E.H., Enns, W.R. & Werner, F.G. (1976) Bionomics of the bee-associated Meloidae (Coleoptera); bee and plant hosts of some Nearctic meloid beetles-a synopsis. Annals of the Entomological Society of America, 69, 959–970.

Erickson, E.H. & Werner, F.G. (1974a) Bionomics of nearctic bee-associated Meloidae (Coleoptera); life histories and nutrition of certain Meloinae. Annals of the Entomological Society of America, 67, 394–400.

Erickson, E.H. & Werner, F.G. (1974b) Bionomics of Nearctic bee-associated Meloidae (Coleoptera); life histories and nutrition of certain Nemognathinae. Annals of the Entomological Society of America, 67, 401–406.

Evans, E.W. (1990) Dynamics of an aggregation of blister beetles (Coleoptera: Meloidae) attacking a prairie legume. Journal of the Kansas Entomological Society, 63, 616–625.

Evans, E.W., Smith, C.C. & Gendron, R.P. (1989) Timing and reproduction in a prairie legume: seasonal impacts of insects and consuming flowers and seeds. Oecologia, 78, 220–230.

Finley, R.W. (1976) Original vegetation cover of Wisconsin, compiled from U.S. General Lands Office notes, provided by Wisconsin Department of Natural Resources. Available from: (accessed 16 October 2013)

Fitch, A. (1859) Third report on the noxious and other insects of the State of New York. New York State Agricultural Society, Albany, New York, 767 pp.

Forbes, S.A. (1880) The food of birds. Bulletin of the Illinois Natural History Survey, 1, 80–148.

Fox, W.B. (1943) Some insects infesting the “selenium indicator” vetches in Saskatchewan. Canadian Entomologist, 75, 206–207.

Frost, C.A. (1912) Note on Tricrania sanguinipennis Say (Coleoptera). Psyche, 19, 208.

Gaston, K.J. (1991) The magnitude of global insect species richness. Conservation Biology, 5, 283–296.

Gibson, A. (1912) Blister beetles. 42nd Annual report of the Entomological Society of Ontario, 36, 83–88.

Gilbertson, G.I. & Horsfall, W.R. (1940) Blister beetles and their control. South Dakota Agricultural Experiment Station Bulletin, 340, 1–23.

Goldburg, R.J. (1987) Sequential flowering of neighboring goldenrods and the movements of the flower predator Epicauta pensylvanica. Oecologia, 74, 247–252.

Goplen, B.P. (1958) Cumino sweet clover attacked by Nuttall’s blister (caragana) beetle. Forage Notes, 4, 23–24.

Gotelli, N.J. (2004) A taxonomic wish-list for community ecology. Philosophical Transactions of the Royal Society of London. Series B, 359, 585–597.

Graenicher, S. (1910) On the habits of beetles of the genus Nemognatha. Entomological News, 21, 72–75.

Groom, M.J. (2006) Threats to biodiversity. In: Groom, M.J., Meffe, G.K. & Carroll, C.R. (Eds.), Principles of Conservation Biology. 3rd Eiditon. Sinauer Associates, Inc. Sunderland, Massachusetts, pp. 63–109.

Harrington, W.H. (1894) Notes on a few Canadian Coleoptera. 25th Annual report of the Entomological Society of Ontario, 47–49.

Harris, T.W. (1841) Report on the insects of Massachusetts, injurious to vegetation. Massachusetts Zoological and Botanical Survey, Cambridge, Massachusetts, 459 pp.

Haskins, M.F., Carrel, J.E., Brill, J., Slagle, A.J. & McCormick, J.P. (1987) Quantitative analysis of cantharidin in juvenile blister beetles (Coleoptera: Meloidae). Transactions of the Missouri Academy of Science, 21, 164.

Horsfall, W.R. (1941) Biology of the black blister beetle (Coleoptera: Meloidae). Annals of the Entomological Society of America, 34, 114–126.

Horsfall, W.R. (1942) Biology of the squash blister beetle. Journal of the Kansas Entomological Society, 15, 93–99.

Horsfall, W.R. (1943) Biology and control of common blister beetles in Arkansas. Arkansas Agricultural Experiment Station Bulletin, 436, 1–55.

Houghton, F. (1947) Say’s blister beetles. Bulletin of the Brooklyn Entomological Society, 42, 103–105.

Howell, M. & Ford, P. (1985). The beetle of Aphrodite. In: The beetle of Aphrodite and other medical mysteries. Random House, New York. pp. 237–267.

Huether, J.P. & Huether, M.K. (2015) Distributional and biological notes on recently collected North American Meloidae (Coleoptera). Pan-Pacific Entomologist, 91, 157–167.

Ingram, J.W. (1927) The striped blister beetle on soybeans. USDA leaflet, 12, 1–5.

Johansen, C. & Mayer, D. (1976) Alkali bees: their biology and management for alfalfa seed production in the Pacific Northwest. Pacific Northwest Extension Publication, PNW 0155, 1–24.

Karras, D.J., Farrell, S.E., Harrigan, R.A., Henretig, F.M. & Gealt, L. (1996) Poisoning from “Spanish fly” (cantharidin). The American Journal of Emergency Medicine, 14, 478–483.

Kerr, J.T. & Parker, L. (1999) The environmental basis of North American species richness patterns among Epicauta (Coleoptera: Meloidae). Biodiversity and Conservation, 8, 617–628.

Kinney, K.K., Peairs, F.B. & Swinker, A.M. (1998) Blister beetles in forage crops. Colorado State University Extension, Publication No. 5.524, 1–3.

Kirk, V.M. & Balsbaugh, E.U. Jr. (1975) A list of the beetles of South Dakota. Agricultural Experiment Station Bulletin, 42, 1–139.

Kirk, K. & Bomar, C.R. (2005) Guide to the grasshoppers of Wisconsin. Wisconsin Department of Natural Resources Publication, PUB-SS-1008 2005, 1–154.

Kriska, N.L. & Young, D.K. (2002) An annotated checklist of Wisconsin Scarabaeoidea (Coleoptera). Insecta Mundi, 16, 31–48.

Krombein, K.V. (1967) Trap-nesting wasps and bees: life histories, nests, and associates. Smithsonian Press, Washington, D.C., 594 pp.

Kumar, R., Lavigne, R.J., Lloyd J.E. & Pfadt, R.E. (1976) Insects of the Central Plains Experiment Range, Pawnee National Grassland. University of Wyoming Science Monograph, 32, 1–62.

Lavigne, R.J. (1976) Rangeland insect-plant association on the Pawnee site. Annals of the Entomological Society of America, 69, 753–763.

Leech, H.B. (1934) Almost a cannibal. Bulletin of the Brooklyn Entomological Society, 29, 41.

LeSage, L. & Bousquet, Y. (1983) A new record of attacks by Pedilus (Pedilidae) on Meloe (Meloidae: Coleoptera). Entomological News, 94, 95–96.

Lisberg, A.E. & Young, D.K. (2003). An annotated checklist of Wisconsin Mordellidae (Coleoptera). Insect Mundi, 17, 195–202.

MacSwain, J.W. (1952) A synopsis of the genus Gnathium, with description of new species (Coleoptera, Meloidae). The Wasmann Journal of Biology, 10, 205–224.

MacSwain, J.W. (1956) A classification of the first instar larvae of the Meloidae (Coleoptera). Annals of the Entomological Society of America, 36, 360–364.

Marschalek, D.A. & Young, D.K. (2013) Mark-recapture study of Gnathium minimum (Say) (Coleoptera: Meloidae) reveals limited dispersal among Helianthus occidentalis patches. Entomological News, 123, 285–291.

May, M.M. (1988) How many species are there on Earth? Science, 241, 1441–1449.

Mayer, D.F. & Johansen, C.A. (1977) Cantharidin from Meloe niger Kirby. Pan-Pacific Entomologist, 53, 101–103.

Mayer, D.F. & Johansen, C.A. (1978) Bionomics of Meloe niger Kirby (Coleoptera: Meloidae), a predator of the alkali bee, Nomia melanderi Cockerall (Hymenoptera: Halictidae). Washington State University, College of Agriculture Research Center, Paper No. 4981, 1–22.

McLain, D.K. (1982) Behavioral and morphological correlates of male dominance and courtship persistence in the blister beetle Epicauta pensylvanica (Coleoptera: Meloidae). American Midland Naturalist, 107, 396–403.

Meyer, D., Schlatter, C., Schlatter-Lanz, I., Schmid, H. & Bovey, P. (1968) Die zucht von Lytta vesicatoria im laboratorium und nachweis der cantharidin-synthese in larven. Experientia, 24, 995–998.

Middlekauff, W.W. (1958) Biology and ecology of several species of California rangeland grasshoppers. Pan-Pacific Entomologist, 34, 1–11.

Mikkelsen, P.M. & Cracraft, J. (2001) Marine biodiversity and the need for systematic inventories. Bulletin of Marine Science, 69, 525–534.

Milliken, F.B. (1921) Results of work on blister beetles in Kansas. USDA Bulletin, 967, 1–26.

Miranpuri, G.S. & Khachatourians, G.G. (1994) Pathogenicity of Beauveria bassiana (Bals.) Vuill. and Verticillium lecannii (Zimm.) toward blister beetle, Lytta nuttali Say (Col., Meloidae). Journal of Applied Entomology, 118, 103–110.

Moore, R.W. (1963) Cantharides poisoning in a horse. Veterinary Medicine, 58, 961.

Munsen, J. (1966) An effective treatment for bilharzial fibrosis and calcification of the bladder. The Journal of Urology, 96, 459–460.

Mutchler, A.J. & Weiss, H.B. (1924) The oil and blister beetles of New Jersey. New Jersey Department of Agriculture Bulletin, 76, 1–19.

Nabli, H., Bailey, W.C. & Necibi, S. (1999) Trapping of Phyllophaga sp. (Coleoptera: Scarabaeidae), Epicauta sp. (Coleoptera: Meloidae), and green stink bugs (Hemiptera: Pentatomidae) with different sources of light. Journal of the Kansas Entomological Society, 72, 222–225.

Nagatomi, A. & Iwata, K. (1958) Biology of the Japanese blister beetle, Epicauta gorhami Marseul. Mushi, 31, 29–47.

Nardi, G. & Bologna, M.A. (2000) Cantharidin attraction in Pyrochroa (Coleoptera: Pyrochroidae). Entomological News, 111, 74–75.

Parker, J.B. & Boving, A.G. (1924) The blister beetle Tricrania sanguinipennis–biology, discriptions of different stages, and systematic relationship. Proceedings of the United States National Museum, 64, 1–40.

Parker, J.R & Wakeland, C. (1957) Grasshopper egg pods destroyed by larvae of bee flies, blister beetles, and ground beetles. USDA Technical Bulletin, 1165, 1–29.

Pechmann, J.H.K. & Wake, D.B. (2006) Enigmatic declines and disappearances of amphibian populations. In: Groom, M.J., Meffe, G.K. & Carroll, C.R. (Eds.), Principles of Conservation Biology. 3rd Edition. Sinauer Associates, Inc. Sunderland, Massachusetts, pp. 93–98.

Pinto, J.D. (1991) The taxonomy of North American Epicauta (Coleoptera: Meloidae), with revision of the nominate subgenus and a survey of courtship behavior. University of California Publication Entomology Bulletin, 110, 1–372.

Pinto, J.D. (2009) A taxonomic review of the genus Gnathium Kirby (Coleoptera: Meloidae). Transactions of the American Entomological Society, 135, 1–58.

Pinto, J.D. & Bologna, M.A. (1999) The new world genera of Meloidae (Coleoptera): a key and synopsis. Journal of Natural History, 33, 569–629.

Pinto, J.D. & Bologna, M.A. (2002) Meloidae. In: Arnett, R.H., Thomas, M.C., Skelley, P.E. & Frank, J.H. (Ed.), American beetles. Vol. II. Polyphaga: Scrabaeoidea through Curculionidea. CRC Press, Boca Raton, Florida, pp. 522–529.

Pinto, J.D., Bologna, M.A. & Bouseman, J.K. (1996) First-instar larvae, courtship and oviposition in Eletica: amending the definition of the Meloidae (Coleoptera: Tenebrionoidea). Systematic Entomology, 21, 63–74.

Pinto, J.D. & Mayor, A.J. (1986) Size, mating success, and courtship pattern in the Meloidae (Coleoptera). Annals of the Entomological Society of America, 79, 597–604.

Pinto, J.D. & Selander, R.B. (1970) The bionomics of blister beetles of the genus Meloe and a classification of the new world species. University of Illinois Press, Urbana Biological Monographs, 42, 1–222.

Pitzalis, M., Amore, V., Montalto, F., Luiselli, L. & Bologna, M.A. (2014) Rarity of blister beetles (Coleoptera: Meloidae) in Southern Africa correlates with their phylogeny and trophic habits, but not body size. European Journal of Entomology, 111, 529–535.

Presto, A.J. III & Muecke, E.C. (1970) A dose of Spanish fly. Journal of the American Medical Association, 214, 591–592.

Price, M.B. & Young, D.K. (2006) An annotated checklist of Wisconsin sap and short-winged flower beetles (Coleoptera: Nitidulidae, Kateretidae). Insecta Mundi, 20, 69–84.

Ray, A.C., Tamulinas, S.H. & Reagor, J.C. (1979) High pressure liquid chromatographic determination of cantharidin, using a derivatilization method in specimens from animals acutely poisoned by ingestion of blister beetles, Epicauta lemniscata. American Journal of Veterinary Research, 40, 498–504.

Ray, A.C., Post, L.O., Hurst, J.M., Edwards, W.C. & Reagor, J.C. (1980) Evaluation of an analytical method for the diagnosis of cantharidin toxicosis due to ingestion of blister beetle (Epicauta lemniscata) by horses and sheep. American Journal of Veterinary Research, 41, 932–933.

Rees, N.E. (1973) Arthropod and nematode parasites, parasitoids, and predators of Acrididae in America north of Mexico. USDA Technical Bulletin, 1460, 1–288.

Riley, C.V. (1877) On the characters and habits of the blister-beetles belonging to the genera Macrobasis Lec. and Epicauta Fabr.; with remarks on other species of the family Meloidae. Transactions of the St. Louis Academy Science, 3, 544–562.

Riley, C.V. (1878) Invertebrate enemies. In: First annual report...for the year 1877 relating to the Rocky Mountain locust and the best methods of preventing its injuries and of guarding against its invasions. United States Entomology Commission Annual Report, pp. 284–334.

Robertson, C. (1928) Flowers and Insects. Science Press Printing Company, Lancaster, Pennsylvania, 221 pp.

Say, T. (1827) Descriptions of new species of coleopterous insects inhabiting the United States. Journal of the Academy of Natural Sciences of Philadelphia, 5, 160–204, 237–284, 293–304.

Schoeb, T.R. & Panciera, R.J. (1978) Blister beetle poisoning in horses. Journal of the American Veterinary Medical Association, 173, 75–77.

Schoeb, T.R. & Panciera, R.J. (1979) Pathology of blister beetle (Epicauta) poisoning in horses. Veterinary Pathology, 16, 18–31.

Schmidt, J.P. (2006) The blister beetles (Coleoptera: Meloidae) of Colorado. Colorado State University Masters of Science Thesis, 235 pp.

Selander, R.B. (1960) Bionomics, systematics, and phylogeny of Lytta, a genus of blister beetles (Coleoptera, Meloidae). Illinois Biological Monographs, 28, 1–295.

Selander, R.B. (1981) Evidence for a third type of larval prey in blister beetles (Coleoptera: Meloidae). Journal of the Kansas Entomological Society, 54, 757–783.

Selander, R.B. (1982) Further studies of predation of meloid eggs by meloid larvae (Coleoptera) Epicauta atrata, North America. Journal of the Kansas Entomological Society, 55, 427–441.

Selander, R.B. (1991) On the nomenclature and classification of the Meloidae (Coleoptera). Insect Mundi, 5, 65–94.

Selander, R.B. & Bohart, G.E. (1954) The biology of Zonitis atripennis flavida LeConte (Coleoptera: Meloidae). The Wasmann Journal of Biology, 12, 227–243.

Selander, R.B. & Fasulo, T.R. (2003) Blister beetles (Insecta: Coleoptera: Meloidae). Entomology and Nematology Department, Cooperative Extension Service, Institute of Food and Agricultural Sciences, University of Florida, Report EENY-166, 1–7.

Selander, R.B. & Mathieu, J.M. (1964) The ontogeny of blister beetles (Coleoptera, Meloidae). I. A study of three species of the genus Pyrota. Annals of the Entomological Society of America, 57, 711–732.

Sherman, F., Jr. (1913) The Meloidae (blister-beetles) of North Carolina (Col.). Entomological News, 24, 245–247.

Sierra, J.R., Woggon, W.-D. & Schmid, H. (1976) Transfer of cantharidin during copulation from the adult male to the female Lytta vesicatoria (‘Spanish flies’). Experientia, 32, 142–144.

Snead, J.S. & Alcock, J. (1985) Aggregation formation and assortative mating in two Meloid beetles. Evolution, 39, 1123–1131.

Staines, C.L. (1983) The Meloidae (Coleoptera) of Maryland. Maryland Entomologist, 2, 41–52.

Swanson, J.I. & Kleintjes Neff, P.K. (2007) Lycaeides melissa samuelis (Lepidoptera: Lycaenidae) response to an aggregation of Lytta sayi (Coleoptera: Meloidae) on Lupinus perennis (Fagaceae). The Great Lakes Entomologist, 40, 69–79.

Thompson, B.T., III. (2004) Wisconsin bird watching: A year-round guide. Cool Springs Press, Nashville, Tennessee, 176 pp.

Ulke, H. (1902) List of beetles of the District of Columbia. Proceedings of the United States National Museum, 25, 1–57.

U. S. Fish & Wildlife Service. (1993) Endangered and threatened wildlife and plants; 90-day finding for a petition to list four California butterflies as endangered and continuation of status reviews. Federal Register, 50 CFR 17, 38549–38552.

U. S. Fish & Wildlife Service. (2006) Endangered and threatened wildlife and plants; 90-day finding for a petition to list the Hermes copper butterfly as endangered. Federal Register, 50 CFR 17, 44966–44976.

Watson, J.R. (1917) Report of the entomologist: Annual report for 1916 and 1917. Florida Agricultural Experiment Station Annual Report, 1917, 52R–65R.

Werner, F.G. (1945) A revision of the genus Epicauta in American north of Mexico. Bulletin of the Museum of Comparative Zoology, 95, 421–517.

Werner, F.G., Enns, W.R. & Parker, F.H. (1966) The Meloidae of Arizona. The University of Arizona Agricultural Experiment Station Technical Bulletin, 175, 1–96.

Williams, A.H. & Young, D.K. (1999) Attraction of Pedilus lugubris (Coleoptera: Pyrochroidae) to Epicauta murina and Epicauta fabricii (Coleoptera: Meloidae) and new food plant records for Epicauta spp. The Great Lakes Entomologist, 32, 97–99.

Wirth, W.W. (1956) The biting midges ectoparasitic on blister beetles (Diptera, Heleidae). Proceedings of the Entomological Society of Washington, 58, 15–23.

Wisconsin Geological Survey. (2012) Early vegetation of Wisconsin map. Wisconsin Geological Survey. Available from: (accessed 16 August 2012)

Young, D.K. (1984a) Cantharidin and insects: an historical review. The Great Lakes Entomologist, 17, 187–194.

Young, D.K. (1984b) Field records and observations of insects associated with cantharidin. The Great Lakes Entomologist, 17, 195–199.

Zhu, Y.-C. & Higgins, R.A. (1994) Host plant influences on feeding, survivorship, population distribution, and management of blister beetles (Coleoptera: Meloidae) in Kansas. Environmental Entomology, 23, 1472–1479.


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