Coleoptera: family-group review and keys to identification

Jan Klimaszewski, J Charles Watt

Abstract


The 82 beetle families represented in New Zealand are reviewed. These families are briefly characterised in the phylogenetic order proposed by Lawrence & Newton (1995)1, with family concepts similar to those defined by these authors and by Pakaluk et al. (1994)2.

A general introduction to the New Zealand beetle fauna is given, and collecting methods are briefly discussed, and core literature is reviewed. Two alternative keys - pyramidal and linear - are provided for identification of the families, using often different character sets. Morphological features are explained, and many are illustrated. Estimated numbers of species, genera, and (where appropriate) tribes and subfamilies are given. Known distributions are discussed, and data on natural history and collecting are summarised. For each family at least one species is represented by a habitus illustration, short description, and geographic profile, and important references are listed.

Keys are provided to subfamilies of Carabidae, Leiodidae, Staphylinidae, Trogossitidae, Phloeostichidae, Scarabaeidae, Cerambycidae, Chrysomelidae, Anthribidae, Belidae, Brentidae, and Curculionidae; for some other families a substantive publication containing a key is cited.

Some 1100 currently recognised genera are recorded for New Zealand, and approximately 5580 species (>5223 native, >356 adventive) are recognised, including undescribed species housed in the New Zealand Arthropod Collection. One endemic family, Chalcodryidae, and four endemic subfamilies - Horelophinae (Hydrophilidae), Euderinae (Bostrichidae), Agapythinae (Phloeostichidae), and Cyclaxirinae (Phalacridae) - are reported. An undescribed species of Platisus from the Three Kings Islands is the only representative of the family Cucujidae in New Zealand.

GENERANATIVE SPECIESADVENTIVE SPECIESSuborder ADEPHAGASuperfamily CARABOIDEA[1] Family Rhysodidae460[2] Family Carabidae (incl. Cicindelinae)75>426>19[3] Family Dytiscidae11160[4] Family Gyrinidae101Suborder POLYPHAGASeries STAPHYLINIFORMIASuperfamily HYDROPHILOIDEA[5] Family Hydrophilidae25705[6] Family Histeridae13236Superfamily STAPHYLINOIDEA[7] Family Hydraenidae5320[8] Family Ptiliidae13>488/9[9] Family Agyrtidae120[10] Family Leiodidae251121[11] Family Scydmaenidae11>2011[12] Family Staphylinidae (incl. Microsilphinae, Pselaphinae, Scaphidiinae)190>936>85Series SCARABAEIFORMIASuperfamily SCARABAEOIDEA[13] Family Lucanidae7263[14] Family Trogidae101[15] Family Scarabaeidae2513212Series ELATERIFORMIASuperfamily SCIRTOIDEA[16] Family Scirtidae (= Helodidae)11>1250[17] Family Eucinetidae110[18] Family Clambidae292Superfamily BUPRESTOIDEA[19] Family Buprestidae221Superfamily BYRRHOIDEA[20] Family Byrrhidae8>790[21] Family Dryopidae240[22] Family Elmidae1160[23] Family Limnichidae271[24] Family Heteroceridae110[25] Family Ptilodactylidae26-80[26] Family Chelonariidae110Superfamily ELATEROIDEA[27] Family Eucnemidae7220[28] Family Elateridae26>1323[29] Family Lycidae101[30] Family Cantharidae3401Series BOSTRICHIFORMIASuperfamily DERODONTOIDEA[31] Family Derodontidae120Superfamily BOSTRICHOIDEA[32] Family Jacobsoniidae23/40[33] Family Nosodendridae12/30[34] Family Dermestidae7116[35] Family Bostrichidae (incl. Lyctinae)417[36] Family Anobiidae (incl. Ptininae)252811Series CUCUJIFORMIASuperfamily CLEROIDEA[37] Family Trogossitidae10>241[38] Family Chaetosomatidae240[39] Family Cleridae8372[40] Family Phycosecidae110[41] Family Melyridae (also as Dasytidae)4330Superfamily CUCUJOIDEA[42] Family Nitidulidae122111[43] Family Monotomidae (incl. Rhizophaginae)215[44] Family Phloeostichidae220[45] Family Silvanidae875[46] Family Cucujidae110[47] Family Laemophloeidae233[48] Family Phalacridae2>31[49] Family Cavognathidae250[50] Family Cryptophagidae121812[51] Family Languriidae580[52] Family Erotylidae290[53] Family Bothrideridae261[54] Family Cerylonidae2>51[55] Family Endomychidae (incl. Merophysinae)361[56] Family Coccinellidae192218[57] Family Corylophidae8>192[58] Family Corticariidae (formerly as Lathridiidae)125311Superfamily TENEBRIONOIDEA[59] Family Mycetophagidae3123[60] Family Archeocrypticidae101[61] Family Ciidae5200[62] Family Melandryidae9380[63] Family Mordellidae461[64] Family Rhipiphoridae350[65] Family Colydiidae26>1960[66] Family Ulodidae (formerly as Zopheridae)4200[67] Family Chalcodryidae350[68] Family Tenebrionidae3613910[69] Family Prostomidae110[70] Family Oedemeridae6183[71] Family Pyrochroidae3>70[72] Family Salpingidae (incl. Inopeplinae)4220[73] Family Anthicidae7179[74] Family Aderidae (formerly as Euglenidae)1/2150[75] Family Scraptiidae2>40Superfamily CHRYSOMELOIDEA[76] Family Cerambycidae56>1808[77] Family Chrysomelidae (incl. Bruchinae)33>134>19Superfamily CURCULIONOIDEA[78] Family Nemonychidae140[79] Family Anthribidae28583[80] Family Belidae (incl. Aglycyderinae = Proterhininae)3110[81] Family Brentidae (incl. Apioninae)331[82] Family Curculionidae (incl. Scolytinae)231>149646Totals (approx.):1100>5223>356

Keywords


Taxonomy; keys; biology; review; Coleoptera; beetles

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