Harpalini (Insecta: Coleoptera: Carabidae: Harpalinae)

André Larochelle, Marie-Claude Larivière

Abstract


The New Zealand Harpalini fauna (Coleoptera: Carabidae: Harpalinae) is revised. Twenty genera and fifty-seven species are recognised. Five new genera are described: Maoriharpalus new genus and Tuiharpalus new genus (Anisodactylina), Hakaharpalus new genus and Kupeharpalus new genus (Pelmatellina), and Kiwiharpalus new genus (Stenolophina). The species Parabaris gourlayi Britton, 1964 is transferred to Tuiharpalus new genus. The original combination Harpalus australasiae Dejean, 1829 is reinstated. Tachys (?) cavelli Broun, 1893 is transferred to the genus Hakaharpalus. Twenty-three new species are described: Anisodactylina — Allocinopus belli new species, A. bousqueti new species, A. wardi new species, Maoriharpalus sutherlandi new species, Parabaris hoarei new species, P. lesagei new species, Tuiharpalus clunieae new species, T. crosbyi new species, T. hallae new species, T. moorei new species; Pelmatellina — Hakaharpalus davidsoni new species, H. maddisoni new species, H. patricki new species, H. rhodeae new species, Kupeharpalus barrattae new species, K. embersoni new species, K. johnsi new species, Lecanomerus marrisi new species, Syllectus gouleti new species; Stenolophina — Kiwiharpalus townsendi new species, Pholeodytes helmorei new species, P. nunni new species, and P. palmai new species . Eight new synonymies are established (valid name listed after equal sign): Allocinopus ocularius Broun, 1908 = Allocinopus sculpticollis Broun, 1903; Allocinopus castaneus Broun, 1912 = Allocinopus smithi Broun, 1912; Hypharpax abstrusus Bates, 1878 = Hypharpax australis (Dejean, 1829); Lecanomerus fallax Broun, 1880 = Lecanomerus insignitus Broun, 1880; Lecanomerus fuliginosus Broun, 1880, Lecanomerus pallipes Broun, 1894, and Lecanomerus incertus Broun, 1914 = Lecanomerus latimanus Bates, 1874; Syllectus spelaeus Britton, 1964 = Syllectus magnus Britton, 1964. Lectotypes are designated for the following taxa: Allocinopus smithi Broun, 1912; A. castaneus Broun, 1912; A. latitarsis Broun, 1911; Euthenarus brevicollis Bates, 1874; E. puncticollis Bates, 1874; Gaioxenus pilipalpis Broun, 1910; Harpalus antarcticus Laporte de Castelnau, 1867, H. novaezelandiae Laporte de Castelnau, 1867, Hypharpax abstrusus Bates, 1878; Lecanomerus fallax Broun, 1880; L. obesulus Bates, 1878; Syllectus anomalus Bates, 1878; Triplosarus fulvescens Bates, 1874. Four adventive species are recorded for New Zealand for the first time: Gnathaphanus melbournensis (Laporte de Castelnau, 1867); Notiobia quadricollis (Chaudoir, 1878); Euthenarus bicolor Moore, 1985; E. promptus (Erichson, 1842).

A concise revision of the taxonomy of all taxa is provided. Subtribes, genera, and species are keyed. Descriptions are provided with illustrations emphasising the most important diagnostic features of the external morphology and male genitalia. Information is given on synonymy, type data, material examined, geographic distribution, ecology, biology, dispersal power, and collecting techniques. The composition of the New Zealand Harpalini fauna, with endemism levels of 55% for genera and 75% for species, and its affinities with Australia, New Caledonia, Lord Howe Island, and Norfolk Island are analysed and discussed. Over 50% of native taxa (25 out of 42 species) are known from 10 populations or fewer and may be of potential conservation concern.

Checklist of taxa

Note: Valid taxa are listed alphabetically in bold italic type; synonyms are listed underneath according to year published in italic type (A=Adventive, E=Endemic, N=Native, but not endemic to New Zealand).

Family CARABIDAE
Subfamily HARPALINAE
Tribe HARPALINI
Subtribe ANISODACTYLINA
Genus Allocinopus Broun, 1903 E
     angustulus Broun, 1912 E
     belli new species E
     bousqueti new species E
     latitarsis Broun, 1911 E
     sculpticollis Broun, 1903 E
          ocularius Broun, 1908 new synonym
     smithi Broun, 1912 E
          castaneus Broun, 1912 new synonym
     wardi new species E

Genus Anisodactylus Dejean, 1829 A
   Subgenus Anisodactylus Dejean, 1829 A
     binotatus (Fabricius, 1787) A

Genus Gaioxenus Broun, 1910 E
     pilipalpis Broun, 1910 E

Genus Gnathaphanus Macleay, 1825 A
         Pachauchenius Macleay, 1864
          Mirosarus Bates, 1878
     melbournensis (Laporte de Castelnau, 1867) A first New Zealand record
          paroensis Laporte de Castelnau, 1867, Harpalus
          marginicollis Laporte de Castelnau, 1867, Harpalus
          adelaideae Laporte de Castelnau, 1867, Harpalus
          planipennis Macleay, 1871, Harpalus
          angustatus Macleay, 1871, Harpalus
          aeneonitens Macleay, 1871, Harpalus
          gayndahensis Macleay, 1871, Harpalus
          atroviridis Macleay, 1871, Harpalus
          insularis Bates, 1878, Mirosarus

Genus Hypharpax Macleay, 1825 N
          Sagraemerus Redtenbacher, 1868
     antarcticus (Laporte de Castelnau, 1867) E
     australis (Dejean, 1829) A
          inornatus Germar, 1848, Harpalus
          coxii Laporte de Castelnau, 1867, Harpalus
          abstrusus Bates, 1878 new synonym
          parvus Chaudoir, 1878

Genus Maoriharpalus new genus E
     sutherlandi new species E

Genus Notiobia Perty, 1830 A
   Subgenus Anisotarsus Chaudoir, 1837 A
          Diaphoromerus Chaudoir, 1843
          Eurytrichus LeConte, 1848
          Stilbolidus Casey, 1914
     quadricollis (Chaudoir, 1878) A first New Zealand record

Genus Parabaris Broun, 1881 E
     atratus Broun, 1881 E
     hoarei new species E
     lesagei new species E

Genus Triplosarus Bates, 1874 E
     novaezelandiae (Laporte de Castelnau, 1867) E
          fulvescens Bates, 1874

Genus Tuiharpalus new genus E
     clunieae new species E
     crosbyi new species E
     gourlayi (Britton, 1964) E new combination
     hallae new species E
     moorei new species E

Subtribe HARPALINA
Genus Harpalus Latreille, 1802 A
   Subgenus Harpalus Latreille, 1802 A
          Amblystus Motschulsky, 1864
     affinis (Schrank, 1781) A
          aeneus Fabricius, 1775, Carabus homonym
     tardus (Panzer, 1797) A

  Subgenus (Uncertain)
     australasiae Dejean, 1829 A reinstated

Subtribe PELMATELLINA
Genus Hakaharpalus new genus E
     cavelli (Broun, 1893) E new combination
     davidsoni new species E
     maddisoni new species E
     patricki new species E
     rhodeae new species E

Genus Kupeharpalus new genus E
     barrattae new species E
     embersoni new species E
     johnsi new species E

Genus Lecanomerus Chaudoir, 1850 N
          Thenarotes Bates,1878
          Odontagonum Darlington, 1956
     atriceps (Macleay, 1871) A
          javanus Jedli…ka, 1964, Acupalpus
     insignitus Broun, 1880 E
          fallax Broun, 1880 new synonym
     latimanus Bates, 1874 E
          fuliginosus Broun, 1880 new synonym
          pallipes Broun, 1894 new synonym
          incertus Broun, 1914 new synonym
     marrisi new species E
     obesulus Bates, 1878 E
     sharpi (Csiki, 1932) E
          marginatus Sharp, 1883 homonym
     verticalis (Erichson, 1842) A
          insidiosus Chaudoir, 1850
          flavocinctus Blackburn, 1888
          occidentalis Sloane, 1898
     vestigialis (Erichson, 1842) A
          mastersii Macleay, 1871, Acupalpus
          stenopus Broun, 1886
          nitidus Blackburn, 1891
          labralis Broun, 1914

Genus Syllectus Bates, 1878 E
     anomalus Bates, 1878 E
     gouleti new species E
     magnus Britton, 1964 E
          spelaeus Britton, 1964 new synonym

Subtribe STENOLOPHINA
Genus Egadroma Motschulsky, 1855 A
     picea (Guérin-Méneville, 1830) A
          dingo Laporte de Castelnau, 1867, Harpalus
          politus Macleay, 1871, Stenolophus
          sexualis Fauvel, 1882, Stenolophus

Genus Euthenarus Bates, 1874 N
     bicolor Moore, 1985 A first New Zealand record
     brevicollis Bates, 1874 E
     promptus (Erichson, 1842) A first New Zealand record
     puncticollis Bates, 1874 E

Genus Haplanister Moore, 1996 A
     crypticus Moore, 1996 A

Genus Kiwiharpalus new genus E
     townsendi new species E

Genus Pholeodytes Britton, 1962 E
     cerberus Britton, 1964 E
     helmorei new species E
     nunni new species E
     palmai new species E
     townsendi Britton, 1962 E


Keywords


Coleoptera; Carabidae; Harpalinae; Harpalini; ground beetles; taxonomy; phylogeny; new genus; new species; biology; morphology

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