Carabidae (Insecta: Coleoptera): synopsis of supraspecific taxa

André Larochelle, Marie-Claude Larivière

Abstract


A synopsis of the New Zealand supraspecific taxa of Carabidae (Coleoptera) and an updated checklist of species-group taxa are presented.

Descriptions (subfamilies to subgenera), identification keys (subfamilies, tribes, and genera), habitus drawings (genera and subgenera), as well as distributional and ecological information, summaries of collecting techniques, and the most relevant bibliographic references for all genera, are provided. An appendix including a checklist of species updating the catalogue of Larochelle & Larivière (2001, Fauna of New Zealand 43) is also provided.

The New Zealand carabid fauna currently includes 7 subfamilies, 21 tribes, 86 genera, and 476 species-group taxa (461 species, plus 15 subspecies). Fifty genera (58%) are currently recognised as being endemic; seventeen genera (20%) are native, and nineteen genera (22%) are adventive.

Three new genera and one new species are described: Kiwitachys new genus (type species: Tachys antarcticus Bates, 1874); Kiwitrechus new genus (type species: Kiwitrechus karenscottae new species); Kupetrechus new genus (type species: Duvaliomimus lamberti Britton, 1960). Three new generic synonymies are made (valid names listed after equal sign): Anomalobroscus Johns, 2007 = Diglymma Sharp, 1886; Taenarthrus Broun, 1914 = Loxomerus Chaudoir, 1842; Zabronothus Broun, 1893 = Cerabilia Laporte de Castelnau, 1867. Anchomenus Bonelli, 1810, is excluded from the New Zealand fauna. “Anchomenus” sensu White, 1846: 3 (and subsequent authors, especially Broun in many papers), nec Bonelli, 1810, is transferred to Ctenognathus Fairmaire, 1843. Twenty-one new combinations are established (valid names listed after equal sign): Anchomenus arnaudensis Broun, 1921 = Ctenognathus arnaudensis (Broun, 1921); Anchomenus colensonis White, 1846 = Ctenognathus colensonis (White, 1846); Anchomenus edwardsii (Bates, 1874) = Ctenognathus edwardsii (Bates, 1874); Anchomenus helmsi Sharp, 1881 = Ctenognathus helmsi (Sharp, 1881); Anchomenus integratus Broun, 1908 = Ctenognathus integratus (Broun, 1908); Anchomenus intermedius Broun, 1908 = Ctenognathus intermedius (Broun, 1908); Anchomenus libitus Broun, 1914 = Ctenognathus libitus (Broun, 1914); Anchomenus macrocoelis Broun, 1908 = Ctenognathus macrocoelis (Broun, 1908); Anchomenus oreobius Broun, 1886 = Ctenognathus oreobius (Broun, 1886); Anchomenus punctulatus Broun, 1877 = Ctenognathus punctulatus (Broun, 1877); Anchomenus sandageri Broun, 1882 = Ctenognathus sandageri (Broun, 1882); Anchomenus sophronitis Broun, 1908 = Ctenognathus sophronitis (Broun, 1908); Anchomenus sulcitarsis Broun, 1880 = Ctenognathus sulcitarsis (Broun, 1880); Anchomenus xanthomelus Broun, 1908 = Ctenognathus xanthomelus (Broun, 1908); Anomalobroscus seclusus Johns, 2007 = Diglymma seclusum (Johns, 2007); Duvaliomimus lamberti Britton, 1960 = Kupetrechus lamberti (Britton, 1960); Tachys antarcticus Bates, 1874 = Kiwitachys antarcticus (Bates, 1874); Tachys latipennis Sharp, 1886 = Kiwitachys latipennis (Sharp, 1886); Taenarthrus philpotti Broun, 1914 = Loxomerus philpotti (Broun, 1914); Zabronothus rufipes Broun, 1893 = Cerabilia rufipes (Broun, 1893); Zabronothus striatulus Broun, 1893 = Cerabilia striatula (Broun, 1893). The following synonymy is reinstated (valid names listed after equal sign): Zolus Sharp, 1886 = Oopterus Guérin-Méneville, 1841. Eight combinations are also reinstated (valid name listed after equal sign): Taenarthrus capito (Jeannel, 1938) = Loxomerus capito Jeannel, 1938; Zolus atratus Broun, 1893 = Oopterus atratus (Broun, 1893); Zolus carinatus (Broun, 1882) = Oopterus carinatus Broun, 1882; Zolus femoralis Broun, 1894 = Oopterus femoralis (Broun, 1894); Zolus helmsi Sharp, 1886 = Oopterus helmsi (Sharp, 1886); Zolus labralis Broun, 1921 = Oopterus labralis (Broun, 1921); Zolus ocularius Broun, 1917 = Oopterus ocularius (Broun, 1917); Zolus subopacus Broun, 1915 = Oopterus subopacus (Broun, 1915). Three adventive taxa are recorded for New Zealand for the first time: Adelotopus macilentus Baehr, 1997; Dromius meridionalis Dejean, 1825; Trigonothops pacifica (Erichson, 1842). The species Notagonum marginellum (Erichson, 1842) is deleted from the New Zealand fauna.

Family Carabidae
I. Subfamily Cicindelinae
1. Tribe Cicindelini
Subtribe Cicindelina
[1] Genus Cicindela Linnaeus, 1758 N
Subgenus Neocicindela Rivalier, 1963 E

II. Subfamily Carabinae
2. Tribe Carabini
Subtribe Carabina
[2] Genus Carabus Linnaeus, 1758 A
Subgenus Archicarabus Seidlitz, 1887 A

3. Tribe Pamborini
[3] Genus Maoripamborus Brookes, 1944 E

III. Subfamily Migadopinae
4. Tribe Amarotypini
[4] Genus Amarotypus Bates, 1872 E

5. Tribe Migadopini
[5] Genus Calathosoma Jeannel, 1938 E
[6] Genus Loxomerus Chaudoir, 1842 E
      Taenarthrus Broun, 1914 new synonym
Subgenus Loxomerus Chaudoir, 1842 E
Subgenus Pristancylus Blanchard, 1853 E

IV. Subfamily Scaritinae
6. Tribe Clivinini
Subtribe Clivinina
[7] Genus Clivina Latreille, 1802 A

V. Subfamily Trechinae
7. Tribe Broscini
Subtribe Creobiina
[8] Genus Bountya Townsend, 1971 E
Subtribe Nothobroscina
[9] Genus Brullea Laporte de Castelnau, 1867 E
[10] Genus Diglymma Sharp, 1886 E
      Anomalobroscus Johns, 2007 new synonym
[11] Genus Mecodema Blanchard, 1843 E
[12] Genus Metaglymma Bates, 1867 E
[13] Genus Oregus Putzeys, 1868 E

8. Tribe Mecyclothoracini
[14] Genus Mecyclothorax Sharp, 1903 N

9. Tribe Meonini
[15] Genus Selenochilus Chaudoir, 1878 E

10. Tribe Tropopterini
[16] Genus Molopsida White, 1846 E

11. Tribe Trechini
Subtribe Aepina
[17] Genus Kenodactylus Broun, 1909 N
[18] Genus Maoritrechus Brookes, 1932 E
Subtribe Trechina
[19] Genus Duvaliomimus Jeannel, 1928 E
[20] Genus Erebotrechus Britton, 1964 E
[21] Genus Kiwitrechus new genus E
[22] Genus Kupetrechus new genus E
[23] Genus Neanops Britton, 1962 E
[24] Genus Scototrechus Britton, 1962 E

12. Tribe Zolini
Subtribe Zolina
Oopterina
[25] Genus Oopterus Guérin-Méneville, 1841 N
      Zolus Sharp, 1886 reinstated synonym
[26] Genus Synteratus Broun, 1909 E

13. Tribe Bembidiini
Subtribe Bembidiina
[27] Genus Bembidion Latreille, 1802 N
Subgenus Ananotaphus Netolitzky, 1931 E
Subgenus Notaphus Stephens, 1827 A
Subgenus Zeactedium Netolitzky, 1931 E
Subgenus Zecillenus Lindroth, 1980 E
Subgenus Zemetallina Lindroth, 1976 E
Subgenus Zeperyphodes Lindroth, 1976 E
Subgenus Zeperyphus Lindroth, 1976 E
Subgenus Zeplataphus Lindroth, 1976 E
Subtribe Tachyina
[28] Genus Kiwitachys new genus E
[29] Genus Paratachys Casey, 1918 A
[30] Genus Pericompsus LeConte, 1852 A
Subgenus Upocompsus Erwin, 1974 A
[31] Genus Polyderis Motschulsky, 1862 A first record
Subtribe Anillina
[32] Genus Hygranillus Moore, 1980 E
[33] Genus Nesamblyops Jeannel, 1937 E
[34] Genus Pelodiaetodes Moore, 1980 E
[35] Genus Pelodiaetus Jeannel, 1937 E
[36] Genus Zeanillus Jeannel, 1937 E

VI. Subfamily Harpalinae
14. Tribe Pterostichini
Subtribe Pterostichina
[37] Genus Aulacopodus Britton, 1940 E
[38] Genus Gourlayia Britton, 1964 E
[39] Genus Holcaspis Chaudoir, 1865 E
[40] Genus Megadromus Motschulsky, 1866 N
Subgenus Megadromus Motschulsky, 1866 E
[41] Genus Neoferonia Britton, 1940 E
[42] Genus Onawea Johns, 2007 E
      “Argutor” sensu Blanchard, 1843, nec Dejean, 1821
[43] Genus Plocamostethus Britton, 1940 E
[44] Genus Prosopogmus Chaudoir, 1865 A
[45] Genus Psegmatopterus Chaudoir, 1878 E
[46] Genus Rhytisternus Chaudoir, 1865 A
[47] Genus Zeopoecilus Sharp, 1886 E

15. Tribe Licinini
Subtribe Dicrochilina
[48] Genus Dicrochile Guérin-Méneville, 1846 N
Subtribe Licinina
[49] Genus Physolaesthus Chaudoir, 1850 N

16. Tribe Harpalini
Subtribe Anisodactylina
[50] Genus Allocinopus Broun, 1903 E
[51] Genus Anisodactylus Dejean, 1829 A
Subgenus Anisodactylus Dejean, 1829 A
[52] Genus Gaioxenus Broun, 1910 E
[53] Genus Gnathaphanus Macleay, 1825 A
[54] Genus Hypharphax Macleay, 1825 N
[55] Genus Maoriharpalus Larochelle & Larivière, 2005 E
[56] Genus Notiobia Perty, 1830 A
Subgenus Anisotarsus Chaudoir, 1837 A
[57] Genus Parabaris Broun, 1881 E
[58] Genus Triplosarus Bates, 1874 E
[59] Genus Tuiharpalus Larochelle & Larivière, 2005 E
Subtribe Harpalina
[60] Genus Harpalus Latreille, 1802 A
Subgenus Harpalus Latreille, 1802 A
Subgenus Uncertain (H. australasiae Dejean, 1829)
Subtribe Pelmatellina
[61] Genus Hakaharpalus Larochelle & Larivière, 2005 E
[62] Genus Kupeharpalus Larochelle & Larivière, 2005 E
[63] Genus Lecanomerus Chaudoir, 1850 N
[64] Genus Syllectus Bates, 1878 E
Subtribe Stenolophina
[65] Genus Egadroma Motschulsky, 1855 A
[66] Genus Euthenarus Bates, 1874 N
[67] Genus Haplanister Moore, 1996 A
[68] Genus Kiwiharpalus Larochelle & Larivière, 2005 E
[69] Genus Pholeodytes Britton, 1962 E

17. Tribe Platynini
Subtribe Sphodrina
[70] Genus Laemostenus Bonelli, 1810 A
Subgenus Laemostenus Bonelli, 1810 A
Subtribe Platynina
[71] Genus Cerabilia Laporte de Castelnau, 1867 E
      Zabronothus Broun, 1893 new synonym
[72] Genus Ctenognathus Fairmaire, 1843 E
      “Anchomenus” sensu White, 1846, nec Bonelli, 1810 new status
[73] Genus Notagonum Darlington, 1952 N
[74] Genus Platynus Bonelli, 1810 N
[75] Genus Prosphodrus Britton, 1959 E

18. Tribe Perigonini
[76] Genus Perigona Laporte de Castelnau, 1835 A
Subgenus Trechicus LeConte, 1853 A

19. Tribe Pentagonicini
[77] Genus Pentagonica Schmidt-Goebel, 1846 N
[78] Genus Scopodes Erichson, 1842 N

20. Tribe Lebiini
Subtribe Pericalina
[79] Genus Agonocheila Chaudoir, 1848 N
[80] Genus Philophlaeus Chaudoir, 1844 A
Subtribe Actenonycina
[81] Genus Actenonyx White, 1846 E
Subtribe Calleidina
[82] Genus Anomotarus Chaudoir, 1875 A
Subgenus Anomotarus Chaudoir, 1875 A
[83] Genus Demetrida White, 1846 N
Subgenus Demetrida White, 1846 E
[84] Genus Trigonothops Macleay, 1864 A
Subgenus Trigonothops Macleay, 1864 A
Subtribe Dromiina
[85] Genus Dromius Bonelli, 1810 A
Subgenus Dromius Bonelli, 1810 A

VII. Subfamily Pseudomorphinae
21. Tribe Pseudomorphini
[86] Genus Adelotopus Hope, 1834 A first record


Keywords


Coleoptera; Carabidae; suprageneric taxa; ground beetles; new genus; new species; Taxonomy; phylogeny; biology; morphology

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* Broun, T. 1902: See Hutton & Broun, 1902.

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